Portal:Amphibians
The Amphibian Portal
Amphibians are ectothermic, anamniotic, four-limbed vertebrate animals that constitute the class Amphibia. In its broadest sense, it is a paraphyletic group encompassing all tetrapods excluding the amniotes (tetrapods with an amniotic membrane, such as modern reptiles, birds and mammals). All extant (living) amphibians belong to the monophyletic subclass Lissamphibia, with three living orders: Anura (frogs and toads), Urodela (salamanders), and Gymnophiona (caecilians). Evolved to be mostly semiaquatic, amphibians have adapted to inhabit a wide variety of habitats, with most species living in freshwater, wetland or terrestrial ecosystems (such as riparian woodland, fossorial and even arboreal habitats). Their life cycle typically starts out as aquatic larvae with gills known as tadpoles, but some species have developed behavioural adaptations to bypass this.
Young amphibians generally undergo metamorphosis from an aquatic larval form with gills to an air-breathing adult form with lungs. Amphibians use their skin as a secondary respiratory interface and some small terrestrial salamanders and frogs lack lungs and rely entirely on their skin. They are superficially similar to reptiles like lizards, but unlike reptiles and other amniotes, require access to water bodies to breed. With their complex reproductive needs and permeable skins, amphibians are often ecological indicators to habitat conditions; in recent decades there has been a dramatic decline in amphibian populations for many species around the globe.
The earliest amphibians evolved in the Devonian period from tetrapodomorph sarcopterygians (lobe-finned fish with articulated limb-like fins) that evolved primitive lungs, which were helpful in adapting to dry land. They diversified and became ecologically dominant during the Carboniferous and Permian periods, but were later displaced in terrestrial environments by early reptiles and basal synapsids (predecessors of mammals). The origin of modern lissamphibians, which first appeared during the Early Triassic, around 250 million years ago, has long been contentious. The most popular hypothesis is that they likely originated from temnospondyls, the most diverse group of prehistoric amphibians, during the Permian period. Another hypothesis is that they emerged from lepospondyls. A fourth group of lissamphibians, the Albanerpetontidae, became extinct around 2 million years ago. (Full article...)
Selected frog article
Pipoidea are a clade of frogs, that contains the most recent common ancestor of living Pipidae and Rhinophrynidae as well as all its descendants. It is broadly equivalent to Xenoanura. (Full article...)
Selected salamander article
The Salamandroidea are a suborder of salamanders, referred to as advanced salamanders. The members of the suborder are found worldwide except for Antarctica, sub-Saharan Africa, and Oceania. They differ from suborder Cryptobranchoidea as the angular and prearticular bones in their lower jaws are fused, their trunk ribs are bicapitate, and all members use internal fertilization. The female is fertilized by means of a spermatophore, a sperm-containing cap placed by the male in her cloaca. The sperm is stored in spermathecae on the roof of the cloaca until it is needed at the time of oviposition.
The earliest known salamandroid fossils remain contested. Some studies suggest that the earliest salamandroids are represented by specimens of the species Beiyanerpeton jianpingensis and Qinglongtriton gangouensis from the Tiaojishan Formation of Inner Mongolia, China, dated to the Late Jurassic about 157 million years ago. Alternative analyses suggest that Beiyanerpeton jianpingensis and Qinglongtriton gangouensis are stem salamanders and that the oldest known certain salamandroid is Valdotriton gracilis from the Early Cretaceous of Spain, about 127 Ma. (Full article...)
List of selected salamander articles
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Did you know? –
- ...that leopard frogs are an environmental indicator species?
- ... that Sri Lanka is home to three endemic frog genera: Adenomus, Nannophrys, and Lankanectes?
- ...that both species of Corroboree frog are critically endangered?
Selected amphibian type
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Caecilians (/sɪˈsɪliən/; New Latin for 'blind ones') are a group of limbless, vermiform (worm-shaped) or serpentine (snake-shaped) amphibians with small or sometimes nonexistent eyes. They mostly live hidden in soil or in streambeds, and this cryptic lifestyle renders caecilians among the least familiar amphibians. Modern caecilians live in the tropics of South and Central America, Africa, and southern Asia. Caecilians feed on small subterranean creatures such as earthworms. The body is cylindrical and often darkly coloured, and the skull is bullet-shaped and strongly built. Caecilian heads have several unique adaptations, including fused cranial and jaw bones, a two-part system of jaw muscles, and a chemosensory tentacle in front of the eye. The skin is slimy and bears ringlike markings or grooves and may contain scales.
Modern caecilians are a clade, the order Gymnophiona /ˌdʒɪmnəˈfaɪənə/ (or Apoda /ˈæpədə/), one of the three living amphibian groups alongside Anura (frogs) and Urodela (salamanders). Gymnophiona is a crown group, encompassing all modern caecilians and all descendants of their last common ancestor. There are more than 220 living species of caecilian classified in 10 families. Gymnophionomorpha is a recently coined name for the corresponding total group which includes Gymnophiona as well as a few extinct stem-group caecilians (extinct amphibians whose closest living relatives are caecilians but are not descended from any caecilian). Some palaeontologists have used the name Gymnophiona for the total group and the old name Apoda for the crown group. However, Apoda has other even older uses, including as the name of a genus of butterfly, making its use potentially confusing and best avoided. 'Gymnophiona' derives from the Greek words γυμνος / gymnos (Ancient Greek for 'naked') and οφις / ophis (Ancient Greek for 'snake'), as the caecilians were originally thought to be related to snakes and to lack scales.
The study of caecilian evolution is complicated by their poor fossil record and specialized anatomy. Genetic evidence and some anatomical details (such as pedicellate teeth) support the idea that frogs, salamanders, and caecilians (collectively known as lissamphibians) are each other's closest relatives. Frogs and salamanders show many similarities to dissorophoids, a group of extinct amphibians in the order Temnospondyli. Caecilians are more controversial; many studies extend dissorophoid ancestry to caecilians. Some studies have instead argued that caecilians descend from extinct lepospondyl or stereospondyl amphibians, contradicting evidence for lissamphibian monophyly (common ancestry). Rare fossils of early gymnophionans such as Eocaecilia and Funcusvermis have helped to test the various conflicting hypotheses for the relationships between caecilians and other living and extinct amphibians. (Full article...)
List of selected amphibian type articles
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Selected images
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Illustration credit: Ernst HaeckelA frog is an amphibian characterized by long hind legs, a short body, webbed digits, protruding eyes and the absence of a tail. Frogs are most noticeable through their call, which can be widely heard during the mating season. It is estimated that up to 20% of amphibian species may care for their young in one way or another, and there is a great diversity of parental behaviours. For example, frogs in the Gastrotheca genus (upper left) carry their eggs in a pouch, and females of the Eleutherodactylus lineatus species (center left) carry their young on their back. -
Photo: Benny TrappAn Italian tree frog (Hyla intermedia) with an inflated vocal sac. Vocal sacs, flexible membranes of skin possessed by most male frogs, are generally used to amplify mating or advertisement calls. However, some species in the family Rhinodermatidae may also use the sac to store or transport their tadpoles. -
Photo credit: Fir0002The Eastern Banjo Frog (Limnodynastes dumerilli) is native to eastern Australia and has been introduced to New Zealand. The frog is also commonly called the pobblebonk after its distinctive "bonk" call, which is likened to a banjo string being plucked. Adults are roughly seven to eight cm long. -
Photo credit: Derek RamseyThe American Green Frog (Rana clamitans) is native to the eastern half of the United States and Canada. Green frogs live wherever there is shallow freshwater: ponds, road-side ditches, lakes, swamps, streams and brooks. The green frog is abundant wherever it occurs. -
Photo credit: Patrick CoinA terrestrial subadult Eastern newt or red eft (Notophthalmus viridescens). Salamanders of the family Salamandridae with aquatic adult stages are called newts. Some newts, including the Eastern newt, have a juvenile terrestrial stage called the eft. The red eft has bright aposematic coloration to warn predators of its highly toxic skin. -
Photograph: Benny TrappThe Epirus water frog (Pelophylax epeiroticus) is a species of frog in the family Ranidae. It is found in western Greece, including Corfu, and the southern areas of Albania. The species is collected from the wild for human consumption. -
Photograph credit: RushenbRaorchestes parvulus is a small bush frog native to tropical southeastern Asia. Although the reproductive strategy of this species has not been studied, members of the family typically make a foam nest while mating in vegetation overhanging a stream. They create the foam by beating their legs, and the eggs are deposited into the nest and covered with seminal fluid before the foam hardens into a protective casing. When the eggs hatch, the young push their way out of the nest and fall into the water below.
This picture shows a male R. parvulus frog, photographed in Phu Kradueng National Park, Thailand, calling by inflating its vocal sac. -
Common toads in amplexusPhoto: Bernie KohlTwo common toads (Bufo bufo) in amplexus, a form of pseudocopulation found in amphibians and limulids. A male common toad grasps a female with his front legs as part of the mating process. Common toads stay in amplexus for several days. As the female lays a long, double string of small black eggs, the male fertilises them with his sperm; the gelatinous egg strings, which may contain 3000 to 6000 eggs and be 3 to 4.5 metres (10 to 15 ft) in length, are later tangled in plant stalks. -
Photo: LokionlyThe Common Indian Toad (Duttaphrynus melanostictus) is native to South Asia. It grows to about 20 cm (8 in) long and breeds during monsoon season. These toads are often seen at night under street lamps especially during times when winged termites swarm. They have been noted to feed on a wide range of invertebrates, including scorpions. -
Photo credit: LiquidGhoulThe Red-eyed Tree Frog (Litoria chloris) is a species of tree frog native to eastern Australia; ranging from north of Sydney to Proserpine in mid-northern Queensland. These frogs typically reach a size of 65 millimetres (2.6 in). Its skin secretions have been found to destroy HIV, without harming healthy T cells. -
Photograph credit: RushenbWallace's flying frog (Rhacophorus nigropalmatus) is a moss frog found in tropical southeastern Asia. It is named after the British naturalist Alfred R. Wallace, who collected the first known specimen of the species. It lives almost exclusively in trees, and when threatened, or in search of prey, will leap from a branch and splay its four webbed feet; the membranes between its toes and the loose skin flaps on its sides catch the air as it falls, helping it to glide. This individual was photographed in Khao Sok National Park, Thailand. -
Photograph: Benny TrappThe Albanian water frog (Pelophylax shqipericus) is a species of true frog in the family Ranidae. It is native to Albania and Montenegro, where it lives in aquatic environments. The frogs are medium-sized. Males sometimes bear a distinctive bright green stripe down the length of the backbone, but otherwise are green to light brown in overall colouring with large black or dark brown spots. Females are olive green or light brown in colour and also bear brown or black large spots. The species is endangered and known populations are currently in decline. -
Photograph: H. KrispThe phantasmal poison frog (Epipedobates tricolor) is a species of poison dart frog endemic to Ecuador. First described by George Albert Boulenger in 1899, this endangered species is found only in rainforests on the slopes of the Andes. -
Photograph credit: Charles James SharpThe marsh frog (Pelophylax ridibundus) is a species of amphibian native to Europe and parts of Asia. It is the largest type of frog in most of its range, growing to a snout-to-vent length of around 100 mm (4 in); tadpoles can reach up to 190 mm (7.5 in) in length, but this usually occurs in places with long winters where the tadpole has time to grow. Marsh frogs hibernate during the winter, either underwater or in burrows, and are able to use the Earth's magnetic field to locate breeding ponds. This marsh frog was photographed in Kampinos National Park, Poland. -
The Blue Mountains Tree Frog (Litoria citropa) is a moderate-sized species of tree frog, up to about 60 mm (2.4 in) in length. It is native to coastal and highland areas of eastern Australia, especially in the Blue Mountains, hence its name.
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Selected toad article
Toad doctors were practitioners of a specific tradition of medicinal folk magic, operating in western England until the end of the 19th century. Their main concern was healing scrofula (then called "the King's Evil," a skin disease), though they were also believed to cure other ailments including those resulting from witchcraft. They cured the sick by placing a live toad, or the leg of one, in a muslin bag and hanging it around the sick person's neck. (Full article...)
Selected caecilian article
The Siphonopidae are the family of common caecilians. They are found in Central and South America. Like other caecilians, they superficially resemble worms or snakes.
They are the sister group to Dermophiidae, also of South America. Siphonopids are oviparous caecilians, meaning they lay eggs. They have imperforated stapes and no inner mandibular teeth. Like species of some other caecilian families, their skulls have relatively few bones, with those present being fused to form a solid ram to aid in burrowing through the soil. The mouth is recessed beneath the snout, and there is no tail. (Full article...)
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