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While you did a really impressive job expanding that article, more than a few of your references are incorrectly formatted, which you can see if you look at the reference section. There are also some missing parameters which makes the references very difficult to interpret. When your readers can't figure out what reference you had in mind, it's little different to the problem created by not having a source at all. Ian (Wiki Ed) (talk) 05:00, 29 September 2015 (UTC)[reply]
Wikipedia editor Animalparty just reviewed your page, and wrote this note for you:
You may want to reduce (or at least explain) the level of jargon and technical terms, see WP:MTAU. Also, it is common (and often preferred, stylistically) to abbreviate the name to P. portalis after first mention.
To reply, leave a comment on Animalparty's talk page.