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Coffea manombensis
Scientific classification
Kingdom:	Plantae
(unranked):	Angiosperms
(unranked):	Eudicots
Order:	Gentianales
Family:	Rubiaceae
Genus:	Coffea
Species:	manombensis
Binomial name
Coffea manombensis

Distribution

Coffea manombensis tree reaches 6-9 meters tall and 5-8 cm in diameter. The bark is a shade of light to dark brown with a fissured longitudinal pattern. The branches tertete an average of 6mm in diameter and could be smooth to longitudinally fissured. They are also light grey, brown, or whitish color. The Branchlets terete are 2.5-4 mm in diameter, and they have rough to smooth with a spongy texture that is peeling. Older trees may have black, light grey, to whitish dots, but younger trees have a very short and erect hairs. Stipule's deltoids have a triangular, 2-2.5 x 2.5 -3(-6) mm, or they have adnate branching along most of it. The tubular, 4 – 5 x 2.5 - 3.4mm, and adnate or free at the base. The falling chartaceous is glabrous and apex to broadly acute, margin glabrous or ciliate and sometimes hyaline. The leaves are ovate to obovate, or elliptic to elliptic-ovate (4 -)6 - 10.2 x (2.2 -)3 - 4.9 cm, subcoriaceous. The petioles are 1.1 - 2 cm long. The base of the leaves attenuates, and margins are subrevolute to flat. The apex is rounded to obtuse. The young plants and seedlings are caudate. The midrib prominent and secondary veins prominent. They have 6 - 8 pairs that ascended at an angle of 30' to 450 which straight to slightly curved. They curvier and join to form a single hooped intramarginal vein, that has a venation prominent beyond it. The reticulate tertiary is higher order venation prominent, and the leaves surface are abaxial and smooth, and the coloration is lighter than the abaxial surface. The venation is also clearer than the abaxial surface. There are usually few domatia present and they indistinct to prominent which is in the axils of the secondary veins, against the midrib. The orifice is very small (0.1-0.2 mm in diameter). The domatia becoming necrotic and the orifice large, 0.5 mm in diameter. The inflorescence stalk is around to 3-8 mm long and composed of a column hidden by calyculi. Each inflorescence has 1 to 4 flowers. The inflorescence axes, pedicels and calyx can be covered with clear exudate. The bud is enclosed by upper calyculus. There are 3 calyculi but can be 2 due to loss of basal or lower calyculus. Cupular are 2 or 4 lobed and glabrous have very sparse instead of thick appressed hairs. The first or basal calyculus is much smaller than the others above it and it is usually broken and not lobed. The middle calyculus is 2 or 4 foliar lobes. The upper calyculus is the largest 4 lobes that often leaf-like. The middle and upper calyculus are covered with colleters at the base with short thick, dark, semi-translucent hairs above the base and on the inside surface of the lobes. The flowers are 5-merous pedicellate and have a smooth texture. All this information is from the research done by Aron P. Davis and Rakotonasolo Franck in their paper on Three New Species of Coffea L. (Rubiaceae) from Madagascar. https://www.jstor.org/stable/4115654?seq=1#metadata_info_tab_contents

Habitat and Ecology

This coffee tree was original found in the Reserve Speciale de Manombo, southern eastern Madagascar, in 1988. The species was easily recognized by it the long corolla tube, prominent venation, and the upper calyculus with large foliar lobes. The leaves are leathery and fleshy, and they mainly show prominently reticulation upon drying. The long corolla tube is unique comparably to other species of Coffea in eastern Madagascar and even in Africa. The uppermost calyculus are well developed and look like elliptic leaves. Another identification would be the very even, truncate calyx-limb, which is seen after the corolla tube has fallen. They are infrequent and highly localized species. They have a small population of only 10 individuals which were found during field work in the Manombo forest reserve.

Morphology

An image of the plant and more detail description can be found in Aron P. Davis’ and Rakotonasolo Franck’s paper, Three New Species of Coffea L. (Rubiaceae) from Madagascar. https://www.jstor.org/stable/4115654?seq=1#metadata_info_tab_contents

Flowers and fruit

There is no description of a fruit from the research given but an image of the flower, leaves, and branches can be seen in figure 1 for document link. https://www.jstor.org/stable/4115654?seq=1#metadata_info_tab_contents

References

Davis, Aaron P., and Franck Rakotonasolo. "Three New Species of Coffea L. (Rubiaceae) from Madagascar." Kew Bulletin 55, no. 2 (2000): 405-16. Accessed April 28, 2021. doi:10.2307/4115654.https://www.jstor.org/stable/4115654?seq=1#metadata_info_tab_contents