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In nitrate poor soils, acidification of the apoplast increases cell wall extensibility and root growth rate. This is believed to be caused by a decrease in nitrate uptake (due to deficit in the soil medium) and supplanted with an increase in chloride uptake. H+ATPase increases the efflux of H+, thus acidifying the apoplast. [1]

  1. ^ Skobelev, O (July 2010). "Accelerated root growth induced by nitrate deficit is related to apoplast acidification". Russian Journal of Plant Physiology. 57 (4): 485–493. Retrieved 24 Feb 2016.