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Nightingale-Thrush relationships

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Does "aurantiirostris-fuscater/mexicanus-dryas" mean that fuscater and mexicanus form a closely related pair, or that aurantiirostris and fuscater form a pair and mexicanus and dryas form another? Whatever the answer is, I think it should be clarified with words rather than punctuation. —JerryFriedman (Talk) 19:52, 14 May 2008 (UTC)[reply]