Sphincterochila

Sphincterochila
	A lateral view of a live individual of Sphincterochila candidissima
	A dorsal view of a live Sphincterochila candidissima
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Order:	Stylommatophora
Infraorder:	Helicoidei
Superfamily:	Helicoidea
Family:	Sphincterochilidae
Genus:	Sphincterochila; Ancey, 1887
Type species
	Helix boissieri; Charpentier, 1847
Synonyms
	Albea Pallary, 1910; Albea (Cariosula) Pallary, 1910; Albea (Sphincterochila) Ancey, 1887; Calcarina Moquin-Tandon, 1848; Candidissima Pallary, 1910; Leucochroa (Calcarina) Moquin-Tandon, 1848; Leucochroa (Sphincterochila) Ancey, 1887; Mima Westerlund, 1886; Rima Pallary, 1910; Sphincterochila (Albea) Pallary, 1910· accepted, alternate representation; Sphincterochila (Cariosula) Pallary, 1910· accepted, alternate representation; Sphincterochila (Cerigottella) E. Gittenberger, 1993· accepted, alternate representation; Sphincterochila (Rima) Pallary, 1910 (junior synonym); Sphincterochila (Sphincterochila) Ancey, 1887 accepted, alternate representation; Sphincterochila (Zilchena) Forcart, 1972· accepted, alternate representation;

Sphincterochila is a genus of air-breathing land snails, terrestrial pulmonate gastropod molluscs in the family Sphincterochilidae.^[2]

Sphincterochila is the type genus of the family Sphincterochilidae.^[3]

Species in the genus Sphincterochila are arid-adapted (adapted to living in very dry conditions).

Species

Species in the genus Sphincterochila include:

Sphincterochila baetica (Rossmässler, 1854)^[4]^[5]
† Sphincterochila baumbergeri (Wenz, 1936)
Sphincterochila candidissima (Draparnaud, 1801)^[6]^[4]^[5]
Sphincterochila cariosa (Olivier, 1801)^[6]^[5] – shell description
Sphincterochila cariosula (Michaud, 1833)^[4]^[5]
Sphincterochila chionudiscus (Boettger, 1874)^[5]
Sphincterochila cyrenaica Sturany, 1908
- Sphincterochila debeauxi elevata Pallary, 1910^[5]
Sphincterochila fimbriata (Bourguignat, 1852)^[5]
Sphincterochila illicita (Mousson, 1874)
Sphincterochila insularis (Boettger, 1894)^[4]
† Sphincterochila jodoti Rey, 1974
Sphincterochila maroccana Pallary, 1917^[5]
- Sphincterochila maroccana conica Pallary, 1926^[5]
- Sphincterochila maroccana depressa Pallary, 1926^[5]
- Sphincterochila maroccana major Pallary, 1926^[5]
- Sphincterochila maroccana minor Pallary, 1926^[5]
- Sphincterochila maroccana tananensis Pallary, 1917^[5]
Sphincterochila mixta Kaltenbach, 1950
Sphincterochila pagodula (Rutland, 1878)^[5]
- Sphincterochila pagodula umbilicata (Ruttland, 1878)^[5]
Sphincterochila pardoi Llabador, 1950^[5]
Sphincterochila prophetarum (Bourguignat, 1852)^[4]^[5]
Sphincterochila tunetana (Pfeiffer, 1850)^[6]
Sphincterochila zonata (Bourguignat, 1853)^[5]^[7]^[8]

Synonyms

Sphincterochila aharonii (Kobelt, 1913):^[5] synonym of Leucochroa aharonii Kobelt, 1913 (taxon inquirendum)
Sphincterochila boissieri (Charpentier, 1847) – type species:^[4] synonym of Sphincterochila zonata zonata (Bourguignat, 1853)
Sphincterochila corrugata Pallary, 1917:^[5] synonym of Albea mayrani var. corrugata Pallary, 1917 (taxon inquirendum)
Sphincterochila debeauxi (Kobelt, 1881):^[5] synonym of Leucochroa debeauxi Kobelt, 1881 (taxon inquirendum)
Sphincterochila marteli (Pallary, 1918):^[5] synonym of Albea marteli Pallary, 1918 (taxon inquirendum)
Sphincterochila mayrani (Gassies, 1856):^[5] synonym of Albea mayrani (Gassies, 1856) (taxon inquirendum)
- Sphincterochila mayrani turrita (Gassies, 1856)^[5]
Sphincterochila roumensiana (Pallary, 1911):^[5] synonym of Albea roumensiana Pallary, 1918 (taxon inquirednum)

Comparison of apertural views of shells of five Sphincterochila species:

References

^ Ancey C. F. (1887). "Description of new genera or sub-genera of Helicidae". (Continued). The Conchologists’ Exchange 2(2): 22-23.
^ ^a ^b MolluscaBase eds. (2020). MolluscaBase. Sphincterochila Ancey, 1887. Accessed through: World Register of Marine Species at: http://www.marinespecies.org/aphia.php?p=taxdetails&id=996739 on 2020-12-05
^ Bouchet, Philippe; Rocroi, Jean-Pierre; Frýda, Jiri; Hausdorf, Bernard; Ponder, Winston; Valdés, Ángel & Warén, Anders (2005). "Classification and nomenclator of gastropod families". Malacologia. 47 (1–2). Hackenheim, Germany: ConchBooks: 1–397. ISBN 3-925919-72-4. ISSN 0076-2997.
^ ^a ^b ^c ^d ^e ^f Species in genus Sphincterochila (n=6). AnimalBase, accessed 10 April 2009.
^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k ^l ^m ⁿ ^o ^p ^q ^r ^s ^t ^u ^v ^w ^x ^y SPHINCTEROCHILIDAE. page 1 and page 2, Femorale, accessed 25 June 2011.
^ ^a ^b ^c Abbes I., Nouira S. & Neubert E. (2011). "Sphincterochilidae from Tunisia, with a note on the subgenus Rima Pallary, 1910 (Gastropoda, Pulmonata)". ZooKeys 151: 1–15. doi:10.3897/zookeys.151.2264.
^ Steinberger Y. (1991). "Biology of Arid region Soils – Faunal Components". In J. Skujins (ed.). Semiarid Lands and Deserts: Soil Resource and Reclamation. Books in soils, plants and the environment. New York: Marcel Dekker, Inc. p. 187. ISBN 0-8247-8388-3.
^ Shachak M.; Orr Y.; Steinberger Y. (1975). "Field observations on the natural history of Sphincterochila (S.) zonata (Bourguignat, 1853) (= S. boissieri Charpentier, 1847)". Argamon: Israel Journal of Malacology. Israel Malacology Society and Municipal Malacology Museum: Nahariya. 5 (1–4): 20–46. ISSN 0334-326X.

External links

Shoshana Goldenberg, Z. Arad; Heller, J. (1989). "Resistance to desiccation and distribution patterns in the land snailSphincterochila". Journal of Zoology. 218 (3): 353. doi:10.1111/j.1469-7998.1989.tb02549.x..
Checa, Antonio G.; Jiménez-Jiménez, Antonio P.; Rivas, Pascual (1998). "Regulation of spiral coiling in the terrestrial gastropod Sphincterochila: An experimental test of the road-holding model". Journal of Morphology. 235 (3): 249–257. doi:10.1002/(SICI)1097-4687(199803)235:3<249::AID-JMOR4>3.0.CO;2-1. PMID 29852723. S2CID 18751120..
Mizrahi, Tal; Heller, Joseph; Goldenberg, Shoshana; Arad, Zeev (2009). "Heat shock proteins and resistance to desiccation in congeneric land snails". Cell Stress and Chaperones. 15 (4): 351–63. doi:10.1007/s12192-009-0150-9. PMC 3082649. PMID 19953352..
http://www.animalbase.uni-goettingen.de/zooweb/servlet/AnimalBase/home/genus?id=678

[1] Ancey C. F. (1887). "Description of new genera or sub-genera of Helicidae". (Continued). The Conchologists’ Exchange 2(2): 22-23.

[WoRMS-2] MolluscaBase eds. (2020). MolluscaBase. Sphincterochila Ancey, 1887. Accessed through: World Register of Marine Species at: http://www.marinespecies.org/aphia.php?p=taxdetails&id=996739 on 2020-12-05

[3] Bouchet, Philippe; Rocroi, Jean-Pierre; Frýda, Jiri; Hausdorf, Bernard; Ponder, Winston; Valdés, Ángel & Warén, Anders (2005). "Classification and nomenclator of gastropod families". Malacologia. 47 (1–2). Hackenheim, Germany: ConchBooks: 1–397. ISBN 3-925919-72-4. ISSN 0076-2997.

[AB_list-4] ^ ^a ^b ^c ^d ^e ^f Species in genus Sphincterochila (n=6). AnimalBase, accessed 10 April 2009.

[Femorale_2011-5] ^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k ^l ^m ⁿ ^o ^p ^q ^r ^s ^t ^u ^v ^w ^x ^y SPHINCTEROCHILIDAE. page 1 and page 2, Femorale, accessed 25 June 2011.

[Abbes_2011-6] Abbes I., Nouira S. & Neubert E. (2011). "Sphincterochilidae from Tunisia, with a note on the subgenus Rima Pallary, 1910 (Gastropoda, Pulmonata)". ZooKeys 151: 1–15. doi:10.3897/zookeys.151.2264.

[Steinberger_1991-7] Steinberger Y. (1991). "Biology of Arid region Soils – Faunal Components". In J. Skujins (ed.). Semiarid Lands and Deserts: Soil Resource and Reclamation. Books in soils, plants and the environment. New York: Marcel Dekker, Inc. p. 187. ISBN 0-8247-8388-3.

[Shachak_1975-8] Shachak M.; Orr Y.; Steinberger Y. (1975). "Field observations on the natural history of Sphincterochila (S.) zonata (Bourguignat, 1853) (= S. boissieri Charpentier, 1847)". Argamon: Israel Journal of Malacology. Israel Malacology Society and Municipal Malacology Museum: Nahariya. 5 (1–4): 20–46. ISSN 0334-326X.

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