Rhamphastosula

Rhamphastosula; Temporal range: early Pliocene; 5–4 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓
	Holotype of Ramphastosula aguirrei, from the Pisco Formation
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Aves
Order:	Suliformes
Family:	Sulidae
Genus:	†Rhamphastosula; Stucchi & Urbina, 2004
Species
	R. ramirezi Stucchi & Urbina, 2004 (type);

Rhamphastosula ("toucan booby") is an extinct genus of sulid bird known from fossils discovered in early Pliocene rocks of Peru. The type and only named species is R. ramirezi, named for Peruvian vertebrate paleontologist Gregorio Ramirez. The genus name is a combination of "Rhamphastos", "toucan" and "sula", an Icelandic word for "fool" that has been used to describe boobies (Sulidae, Sula) in general.^[1]

Rhamphastosula is known from a partial skull that has bony characters placing it in the booby family Sulidae. Unlike living boobies, it has a large bill with a convex upper mandible, reminiscent of a toucan. The remains were recovered from the west side of Sud Sacaco Level (4–5 Million year-old) of the Pisco Formation. This area was a sheltered beach with rock platforms.^[1]

References

^ ^a ^b Stucchi, Marcelo; Urbina, Mario (2004). ""Ramphastosula" (Aves, Sulidae): A New Genus from the Early Pliocene of the Pisco Formation, Peru". Journal of Vertebrate Paleontology. 24 (4): 974–978. doi:10.1671/0272-4634(2004)024[0974:RASANG]2.0.CO;2. JSTOR 4524793.

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[stucchi04-1] Stucchi, Marcelo; Urbina, Mario (2004). ""Ramphastosula" (Aves, Sulidae): A New Genus from the Early Pliocene of the Pisco Formation, Peru". Journal of Vertebrate Paleontology. 24 (4): 974–978. doi:10.1671/0272-4634(2004)024[0974:RASANG]2.0.CO;2. JSTOR 4524793.

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