2020 in paleobotany

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This article records new taxa of fossil plants that are scheduled to be described during the year 2020, as well as other significant discoveries and events related to paleobotany that are scheduled to occur in the year 2020.

Flowering plants

Alismatales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Limnobiophyllum pedunculatum^[1]	Sp. nov	Valid	Low, Su & Xing in Low et al.	Late Oligocene		China	A member of the family Araceae.

Apiales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Paleopanax puryearensis^[2]	Sp. nov	Valid	Na, Blanchard & Wang	Middle Eocene	Cockfield Formation	United States ( Tennessee)	A member of the family Araliaceae.

Arecales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Echimonocolpites chicxulubensis^[3]	Sp. nov	Valid	Smith et al.	Eocene (Ypresian)		Mexico	Pollen of a flowering plant, probably a member of the family Arecaceae.
Palmoxylon ceroxyloides^[4]	Sp. nov	In press	Khan, Hazra & Bera in Khan et al.	Late Cretaceous (Maastrichtian)-Paleocene (Danian)	Deccan Intertrappean Beds	India	A petrified palm stem of a member of the subfamily Ceroxyloideae.
Palmoxylon dindoriensis^[5]	Sp. nov	Valid	Khan, Roy & Bera in Khan et al.	Late Cretaceous (Maastrichtian)-Paleocene (Danian)	Deccan Intertrappean Beds	India	A petrified palm stem.
Sabalites dawsonii^[6]	Sp. nov	Valid	Greenwood & Conran	Eocene		Canada ( British Columbia)
Sabalites karondiensis^[7]	Sp. nov	In press	Roy, Hazra & Khan in Roy et al.	Late Cretaceous-Paleocene (latest Maastrichtian-earliest Danian)	Deccan Intertrappean Beds	India	A palm frond .
Spinizonocolpites riochiquensis^[8]	Sp. nov	Valid	Vallati & De Sosa Tomas in Vallati, De Sosa Tomas & Casal	Late Cretaceous (Maastrichtian)	Lago Colhué Huapí Formation	Argentina	A member of Arecaceae described on the basis of fossil pollen grains. Announced in 2019; the final version of the article naming it was published in 2020.

Buxales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Pachysandra europaea^[9]	Sp. nov	Valid	Kvaček, Teodoridis & Denk	Pliocene		Germany	A species of Pachysandra. Announced in 2019; the final version of the article naming it was published in 2020.

Caryophyllales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Gomphrenipollis garciae^[10]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin	Brazil	Pollen of a flowering plant, possibly produced by members of the family Amaranthaceae. Announced in 2020; the final version of the article naming it was published in 2021.

Chloranthales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Sarcandraxylon^[11]	Gen. et sp. nov	Valid	Pipo, Iglesias & Bodnar	Late Cretaceous (early–middle Campanian)	Santa Marta Formation	Antarctica	A member of the family Chloranthaceae. Genus includes new species S. sanjosense.

Cornales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Amersinia littletonensis^[12]	Sp. nov	Valid	Huegele & Manchester	Early Paleocene	Denver Formation	United States ( Colorado)
Langtonia parva^[12]	Sp. nov	Valid	Huegele & Manchester	Early Paleocene	Denver Formation	United States ( Colorado)	A member of the family Mastixiaceae.
Mastixicarpum hoodii^[12]	Sp. nov	Valid	Huegele & Manchester	Early Paleocene	Denver Formation	United States ( Colorado)	A member of the family Mastixiaceae.
Nyssa gergoei^[13]	Sp. nov	Valid	Hably	Miocene		Hungary	A tupelo.
Nyssa gyoergyi^[13]	Sp. nov	Valid	Hably	Miocene		Hungary	A tupelo.
Platycrater iljinskajae^[14]	Sp. nov	In press	Denk et al.	Late Oligocene		Russia
Portnallia alexanderi^[12]	Sp. nov	Valid	Huegele & Manchester	Early Paleocene	Denver Formation	United States ( Colorado)	A member of the family Mastixiaceae.

Crossosomatales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Staphylea woodworthensis^[15]	Sp. nov	In press	Zhu & Manchester	Oligocene	Renova Formation	United States ( Montana)	A species of Staphylea.

Cucurbitales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Coriaripites goodii^[16]	Sp. nov	Valid	Barreda, Palazzesi & Tellería in Renner et al.	Late Cretaceous (Campanian–early Maastrichtian)	Lopez de Bertodano Formation Santa Marta Formation Snow Hill Island Formation	Antarctica	Pollen grains similar to those of extant members of the genus Coriaria.
Echitriporites jolyi^[10]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin	Brazil	Pollen produced by members of the genus Cayaponia. Announced in 2020; the final version of the article naming it was published in 2021.

Ericales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Andrewsiocarpon puryearensis^[2]	Sp. nov	Valid	Na, Blanchard & Wang	Middle Eocene	Cockfield Formation	United States ( Tennessee)	A member of the family Theaceae.
Anubiscarpon^[17]	Gen. et sp. nov	Valid	Smith & Manchester	Middle Eocene	Clarno Formation	United States ( Oregon)	A member of the family Theaceae. Genus includes new species A. andersonae.

Fabales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Cercis zekuensis^[18]	Sp. nov	Valid	Li et al.	Early Miocene		China	A species of Cercis. Announced in 2020; the final version of the article naming it was published in 2021.
Gleditsia pliocaenica^[9]	Sp. nov	Valid	Kvaček, Teodoridis & Denk	Pliocene		Germany	A species of Gleditsia. Announced in 2019; the final version of the article naming it was published in 2020.
Menendoxylon lutzi^[19]	Sp. nov	In press	Baez & Crisafulli	Miocene	Chiquimil	Argentina	Fossil wood of a member of the family Fabaceae.
Parkiidites marileae^[10]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin	Brazil	Pollen produced by members of the genus Parkia. Announced in 2020; the final version of the article naming it was published in 2021.
Prioria martineziorum^[20]	Sp. nov	Valid	Rodríguez-Reyes & Estrada-Ruiz	Oligocene-Miocene	Santiago Formation	Panama	A species of Prioria.
Psilastephanocolporites deoliverae^[10]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin	Brazil	Pollen produced by members of the family Polygalaceae. Announced in 2020; the final version of the article naming it was published in 2021.
Psilastephanocolporites endoporatus^[10]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin	Brazil	Announced in 2020; the final version of the article naming it was published in 2021.
Striatopollis grahamii^[3]	Sp. nov	Valid	Smith et al.	Eocene (Ypresian)		Mexico	Pollen of an eudicot, probably a member of the family Fabaceae.

Fagales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Alnus chaybulakensis^[21]	Sp. nov	Valid	Averyanova & Xing	Paleogene		Kazakhstan	An alder.
Berryophyllum hainanensis^[22]	Sp. nov	Valid	Liu & Jin in Liu, Song & Jin	Eocene	Changchang Formation	China	A member of the family Fagaceae.
Carpinus asymmetrica^[23]	Sp. nov	In press	Xue & Jia in Xue et al.	Early Miocene	Maguan	China	A species of Carpinus.
Carpinus symmetrica^[23]	Sp. nov	In press	Xue & Jia in Xue et al.	Early Miocene	Maguan Basin	China	A species of Carpinus
Carya pipecreekensis^[24]	Sp. nov	In press	Swinehart & Farlow	Late Neogene	Pipe Creek Sinkhole	United States ( Indiana)	A hickory.
Castaneophyllum hainanensis^[22]	Sp. nov	Valid	Liu & Jin in Liu, Song & Jin	Eocene	Changchang Formation	China	A member of the family Fagaceae.
Castaneophyllum lanceolata^[22]	Sp. nov	Valid	Liu & Jin in Liu, Song & Jin	Eocene	Changchang Formation	China	A member of the family Fagaceae.
Castanopsis bulgarica^[25]	Sp. nov	Valid	Mantzouka, Ivanov & Bozukov	Late Miocene–early Pliocene (late Messinian–early Zanclean)	Pokrovnik	Bulgaria	A species of Castanopsis. Announced in 2020; the final version of the article naming it was published in 2021.
Lithocarpus changchangensis^[22]	Sp. nov	Valid	Liu & Jin in Liu, Song & Jin	Eocene	Changchang Formation	China	A species of Lithocarpus.
Quercus aimeeana^[26]	Sp. nov	Valid	Sadowski, Schmidt & Denk	Eocene	Baltic amber	Europe (Baltic Sea region)	An oak.
Quercus borissovii^[21]	Sp. nov	Valid	Averyanova & Xing	Paleogene		Kazakhstan	An oak.
Quercus campanulata^[26]	Sp. nov	Valid	Sadowski, Schmidt & Denk	Eocene	Baltic amber	Europe (Baltic Sea region)	An oak.
Quercus casparyi^[26]	Sp. nov	Valid	Sadowski, Schmidt & Denk	Eocene	Baltic amber	Europe (Baltic Sea region)	An oak.
Quercus changchangensis^[22]	Sp. nov	Valid	Liu & Jin in Liu, Song & Jin	Eocene	Changchang Formation	China	An oak.
Quercus conwentzii^[26]	Sp. nov	Valid	Sadowski, Schmidt & Denk	Eocene	Baltic amber	Europe (Baltic Sea region)	An oak.
Quercus longianthera^[26]	Sp. nov	Valid	Sadowski, Schmidt & Denk	Eocene	Baltic amber	Europe (Baltic Sea region)	An oak.
Quercus multipilosa^[26]	Sp. nov	Valid	Sadowski, Schmidt & Denk	Eocene	Baltic amber	Europe (Baltic Sea region)	An oak.
Quercus paleoargyrotricha^[22]	Sp. nov	Valid	Liu & Jin in Liu, Song & Jin	Eocene	Changchang Formation	China	An oak.
Quercus paleohypargyrea^[22]	Sp. nov	Valid	Liu & Jin in Liu, Song & Jin	Eocene	Changchang Formation	China	An oak.
Quercus paleolamellosa^[22]	Sp. nov	Valid	Liu & Jin in Liu, Song & Jin	Eocene	Changchang Formation	China	An oak.

Garryales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Eucommia szaferi^[9]	Sp. nov	Valid	Kvaček, Teodoridis & Denk	Pliocene		Germany	A species of Eucommia. Announced in 2019; the final version of the article naming it was published in 2020.

Gentianales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Asclepiadospermum^[27]	Gen. et 2 sp. nov	Valid	Del Rio et al.	Early Eocene	Niubao Formation	China	An asclepiadoid Apocynaceae genus. Included species A. marginatum and A. ellipticum.
Margocolporites carinae^[10]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin	Brazil	Pollen produced by members of the genus Rauvolfia. Announced in 2020; the final version of the article naming it was published in 2021.

Icacinales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Iodes elliptica^[28]	Sp. nov	In press	Del Rio et al.	Early Oligocene	Wenshan Basin	China	A member of the family Icacinaceae.
Iodes passiciensis^[29]	Sp. nov	Valid	Del Rio & De Franceschi	Early Eocene		France	A member of the family Icacinaceae.
Manchesteria^[30]	Gen. et sp. nov	In press	Stull & Rozefelds in Rozefelds et al.	Cenozoic (mid-Miocene or, more likely, middle Eocene)		Australia	A member of the family Icacinaceae. Genus includes new species M. australis.
Pyrenacantha simonsii^[31]	Sp. nov	Valid	Stull et al.	Early Oligocene		Egypt	A species of Pyrenacantha.

Lamiales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Echitricolpites cruziae^[10]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin	Brazil	Pollen produced by members of the genus Aegiphila. Announced in 2020; the final version of the article naming it was published in 2021.
Multiareolites? reticulatus^[10]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin	Brazil	Pollen produced by members of the family Acanthaceae. Announced in 2020; the final version of the article naming it was published in 2021.
Retistephanocolpites curvimuratus^[10]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin	Brazil	Announced in 2020; the final version of the article naming it was published in 2021.
Retistephanocolpites pardoi^[10]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin	Brazil	Pollen of a flowering plant, possibly produced by members of the genus Amphilophium. Announced in 2020; the final version of the article naming it was published in 2021.

Laurales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Actinodaphnoxylon^[32]	Gen. et sp. nov	In press	Akkemik et al.	Eocene (Lutetian)		Turkey	A member of the family Lauraceae. Genus includes new species A. zileensis.
Mezilaurinoxylon oleiferum^[33]	Sp. nov	Valid	Ruiz, Brea & Pujana in Ruiz et al.	Paleocene (Danian)	Salamanca Formation	Argentina	A member of the family Lauraceae. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.
Patagonoxylon^[33]	Gen. et sp. nov	Valid	Ruiz, Brea & Pujana in Ruiz et al.	Paleocene (Danian)	Salamanca Formation	Argentina	A member of Laurales of uncertain phylogenetic placement. Genus includes new species P. scalariforme. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.
Thymolepis^[34]	Gen. et sp. nov	Valid	Chambers & Poinar	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	Possibly an early representative of Monimiaceae. Genus includes new species T. toxandra.
Valviloculus^[35]	Gen. et sp. nov	Valid	Poinar et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	Possibly a member of Laurales related to the families Monimiaceae and Atherospermataceae. Genus includes new species V. pleristaminis.

Liliales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Smilax fujianensis^[36]	Sp. nov	Valid	Dong et al.	Middle Miocene		China	A species of Smilax. Announced in 2020; the final version of the article naming it was published in 2021.
Smilax zhangpuensis^[36]	Sp. nov	Valid	Dong et al.	Middle Miocene		China	A species of Smilax. Announced in 2020; the final version of the article naming it was published in 2021.

Magnoliales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Magnolia nanningensis^[37]	Sp. nov	Valid	Huang et al.	Late Oligocene	Nanning Basin	China	A species of Magnolia.
Magnolia waltheri^[9]	Sp. nov	Valid	Kvaček, Teodoridis & Denk	Pliocene		Germany	A species of Magnolia. Announced in 2019; the final version of the article naming it was published in 2020.
Melloniflora^[38]	Gen. et sp. nov	Valid	Friis, Crane & Pedersen	Early Cretaceous	Potomac Group	United States ( Virginia)	A relative of extant early-diverging members of the Magnoliales. Genus includes new species M. virginiensis.

Malpighiales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Salix palaeofutura^[39]	Sp. nov	Valid	Narita et al.	Miocene	Bifuka Formation	Japan	A willow.

Malvales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Bastardioxylon^[19]	Gen. et sp. nov	In press	Baez & Crisafulli	Miocene	Chiquimil	Argentina	Fossil wood of a member of the family Malvaceae. Genus includes new species B. antiqua.
Dipterocarpus dindoriensis^[40]	Sp. nov	Valid	Khan, Spicer & Bera in Khan et al.	Late Cretaceous (Maastrichtian)	Deccan Intertrappean Beds	India	A species of Dipterocarpus.
Echiperiporites germeraadii^[10]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin	Brazil	Announced in 2020; the final version of the article naming it was published in 2021.
Echiperiporites jaramilloi^[10]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin	Brazil	Pollen produced by members of the genus Hibiscus. Announced in 2020; the final version of the article naming it was published in 2021.
Echiperiporites titanicus^[10]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin	Brazil	Pollen produced by members of the genus Malachra. Announced in 2020; the final version of the article naming it was published in 2021.
Retistephanocolporites elizabeteae^[10]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin	Brazil	Pollen produced by members of the genus Ceiba. Announced in 2020; the final version of the article naming it was published in 2021.
Veraguasoxylon^[20]	Gen. et sp. nov	Valid	Rodríguez-Reyes & Estrada-Ruiz	Oligocene-Miocene	Santiago Formation	Panama	A member of the family Malvaceae. Genus includes new species V. panamense.

Myrtales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Eucalyptus xoshemium^[41]	Sp. nov	Valid	Gandolfo & Zamaloa in Zamaloa, Gandolfo & Nixon	Eocene (Ypresian)	Huitrera Formation	Argentina	A species of Eucalyptus.
Mangroveoxylon^[42]	Gen. et comb. nov	In press	Moya & Brea	Late Miocene?	Ituzaingó Formation	Argentina	A member of the family Combretaceae; a new genus for "Menendoxylon" areniensis Lutz (1979).
Myrceugenellites grandiporosum^[33]	Sp. nov	Valid	Ruiz, Brea & Pujana in Ruiz et al.	Paleocene (Danian)	Salamanca Formation	Argentina	A member of the family Myrtaceae. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.
Primotrapa^[43]	Gen. et sp. nov	Valid	Li & Li in Li et al.	Late Eocene to early Miocene	Hannuoba	China Czech Republic France Germany	A member of Trapoideae. Genus includes new species P. weichangensis, as well as "Carpolithus" pomelii Saporta (1878) and "Hemitrapa" alpina Su & Zhou in Su et al. (2018).

Nymphaeales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Praenymphaeapollenites^[44]	Gen. et sp. nov	Valid	Barrón, Peris & Labandeira in Peris et al.	Cenomanian	Burmese amber	Myanmar	Pollen of a member of Nymphaeaceae. Genus includes new species P. cenomaniensis.

Oxalidales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Cunoniantha^[45]	Gen. et sp. nov	Valid	Jud & Gandolfo	Paleocene (early Danian)	Salamanca Formation	Argentina	A member of the family Cunoniaceae. Genus includes new species C. bicarpellata. Announced in 2020; the final version of the article naming it was published in 2021.
Elaeocarpus nanningensis^[46]	Sp. nov	In press	Liu et al.	Late Oligocene	Yongning Formation	China	A species of Elaeocarpus.
Elaeocarpus prelacunosus^[46]	Sp. nov	In press	Liu et al.	Late Miocene	Foluo Formation	China	A species of Elaeocarpus.
Elaeocarpus preprunifolioides^[46]	Sp. nov	In press	Liu et al.	Late Miocene	Foluo Formation	China	A species of Elaeocarpus.
Elaeocarpus prerugosus^[46]	Sp. nov	In press	Liu et al.	Late Miocene	Foluo Formation	China	A species of Elaeocarpus.
Elaeocarpus preserratus^[46]	Sp. nov	In press	Liu et al.	Late Miocene	Foluo Formation	China	A species of Elaeocarpus.
Elaeocarpus presikkimensis^[46]	Sp. nov	In press	Liu et al.	Miocene	Erzitang Formation	China	A species of Elaeocarpus.

Poales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Monoporopollenites scabratus^[10]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin	Brazil	Pollen produced by members of the family Poaceae. Announced in 2020; the final version of the article naming it was published in 2021.
Rhizomatites^[47]	Gen. et sp. nov	In press	Robledo & Anzótegui in Robledo et al.	Miocene-Pliocene		Argentina	A member of Cyperaceae. Genus includes new species R. cyperoides.

Proteales

Name	Novelty	Status	Authors	Age	Location	Notes
Banksia microphylla^[48]	Sp. nov	Valid	Carpenter in Carpenter & Milne	Late Eocene	Australia	A species of Banksia.
Banksieaeidites zanthus^[48]	Sp. nov	Valid	Milne in Carpenter & Milne	Late Eocene	Australia	A Banksia-like pollen.
Platanus emryi^[49]	Sp. nov	Valid	Huegele, Spielbauer & Manchester	Miocene	United States	A species of Platanus.

Ranunculales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Cissampelos defranceschii^[50]	Sp. nov	Valid	Del Rio & Su in Del Rio et al.	Middle Eocene	Niubao Formation	China	A species of Cissampelos. Announced in 2020; the final version of the article naming it was published in 2021.
Clematis csabae^[13]	Sp. nov	Valid	Hably	Miocene		Hungary	A species of Clematis.
Diploclisia praeaffinis^[51]	Sp. nov	Valid	Jia et al.	Late Miocene		China	A member of the family Menispermaceae. Announced in 2020; the final version of the article naming it was published in 2021.
Menispermites bangorensis^[50]	Sp. nov	Valid	Huang in Del Rio et al.	Middle Eocene	Niubao Formation	China	A member of the family Menispermaceae. Announced in 2020; the final version of the article naming it was published in 2021.
Menispermites haominae^[50]	Sp. nov	Valid	Huang in Del Rio et al.	Middle Eocene	Niubao Formation	China	A member of the family Menispermaceae. Announced in 2020; the final version of the article naming it was published in 2021.
Menispermites tibetica^[50]	Sp. nov	Valid	Huang in Del Rio et al.	Middle Eocene	Niubao Formation	China	A member of the family Menispermaceae. Announced in 2020; the final version of the article naming it was published in 2021.
Paleoorbicarpum^[52]	Gen. et sp. nov	Valid	Han et al.	Paleocene	Sanshui Basin	China	A member of the family Menispermaceae. Genus includes new species P. parvum.
Stephania bangorensis^[50]	Sp. nov	Valid	Del Rio & Su in Del Rio et al.	Middle Eocene	Niubao Formation	China	A species of Stephania. Announced in 2020; the final version of the article naming it was published in 2021.
Stephania geniculata^[52]	Sp. nov	Valid	Han et al.	Paleocene	Sanshui Basin	China	A species of Stephania
Stephania ornamenta^[52]	Sp. nov	Valid	Han et al.	Paleocene	Sanshui Basin	China	A species of Stephania
Stephania shuangxingii^[50]	Sp. nov	Valid	Del Rio & Su in Del Rio et al.	Middle Eocene	Niubao Formation	China	A species of Stephania. Announced in 2020; the final version of the article naming it was published in 2021.

Rosales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Berhamniphyllum junrongii^[53]	Sp. nov	Valid	Zhou, Wang & Huang in Zhou et al.	Late Eocene	Markam Basin	China	A member of the family Rhamnaceae
Crataegus pentagynoides^[9]	Sp. nov	Valid	Kvaček, Teodoridis & Denk	Pliocene		Germany	A species of Crataegus. Announced in 2019; the final version of the article naming it was published in 2020.
Hemiptelea kryshtofovichii^[21]	Sp. nov	Valid	Averyanova & Xing	Paleogene		Kazakhstan	A member of the family Ulmaceae.
Scabrastephanoporites^[3]	Gen. et sp. nov	Valid	Smith et al.	Eocene (Ypresian)		Mexico	Pollen of an eudicot, probably a member of the family Ulmaceae or Cannabaceae. Genus includes new species S. variabilis.

Sapindales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Acer dombeyopsis^[9]	Sp. nov	Valid	Kvaček, Teodoridis & Denk	Pliocene		Germany	A maple. Announced in 2019; the final version of the article naming it was published in 2020.
Acer viburnoides^[9]	Sp. nov	Valid	Kvaček, Teodoridis & Denk	Pliocene		Germany	A maple. Announced in 2019; the final version of the article naming it was published in 2020.
Acer vitiforme^[9]	Sp. nov	Valid	Kvaček, Teodoridis & Denk	Pliocene		Germany	A maple. Announced in 2019; the final version of the article naming it was published in 2020.
Brosipollis reticulatus^[3]	Sp. nov	Valid	Smith et al.	Eocene (Ypresian)		Mexico	Pollen of a flowering plant, probably a member of the family Burseraceae.
Choerospondias fujianensis^[54]	Sp. nov	In press	Wang et al.	Miocene		China	A species of Choerospondias.
Llanodelacruzoxylon^[55]	Gen. et sp. nov		Rodríguez-Reyes, Estrada-Ruiz & Gasson	Oligocene–Miocene	Santiago Formation	Panama	A member of the family Anacardiaceae. Genus includes new species L. sandovalii.
Manchestercarpa^[56]	Gen. et sp. nov	Valid	Atkinson	Late Cretaceous (Campanian)		Canada ( British Columbia)	A member of the family Meliaceae described on the basis of a fossil fruit. Genus includes new species M. vancouverensis.
Parametopioxylon^[57]	Gen. et sp. nov	Valid	Franco et al.	Miocene	Ituzaingó Formation	Argentina	A member of the family Anacardiaceae described on the basis of fossil wood. Genus includes new species P. crystalliferum.
Psilastephanocolporites hammenii^[3]	Sp. nov	Valid	Smith et al.	Eocene (Ypresian)		Mexico	Pollen of a flowering plant, probably a member of the family Meliaceae
Quinquala^[58]	Gen. et sp. nov	Valid	Manchester & Disney in Manchester, Disney & Pham	Eocene	Clarno Formation Tepee Trail Formation	United States ( Oregon Wyoming)	A fossil fruit with affinities with the Rutaceae. Genus includes new species Q. obovata.
Rousea cavitata^[10]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin	Brazil	Announced in 2020; the final version of the article naming it was published in 2021.

Saxifragales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Corylopsis grisea^[59]	Sp. nov	Valid	Quirk & Hermsen	Early Pliocene	Gray Fossil Site	United States ( Tennessee)	A species of Corylopsis. Announced in 2020; the final version of the article naming it was published in 2021.
Itea polyneura^[60]	Sp. nov	In press	Huang & Tian in Tian et al.	Oligocene	Huazhige Formation	China	A species of Itea.
Protoaltingia^[61]	Gen. et sp. nov	Valid	Scharfstein, Stockey & Rothwell	Late Cretaceous (Coniacian)		Canada ( British Columbia)	A member of the family Altingiaceae. Genus includes new species P. comoxense.

Solanales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Physalis hunickenii^[62]	Sp. nov	Valid	Deanna, Wilf & Gandolfo	Early Eocene	Laguna del Hunco Formation	Argentina	A species of Physalis.

Trochodendrales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Eotrochion^[63]	Gen. et sp. nov	Valid	Manchester, Kvaček & Judd	Paleocene		United States ( Wyoming)	A member of the family Trochodendraceae. Genus includes new species E. polystylum. Announced in 2020; the final version of the article naming it was published in 2021.
Paraconcavistylon^[63]	Gen. et comb. nov	Valid	Manchester, Kvaček & Judd	Ypresian	Klondike Mountain Formation	United States ( Washington)	A Trochodendraceae; a new genus for "Concavistylon" wehrii Manchester et al. (2018). Announced in 2020; the final version of the article naming it was published in 2021.
Trochodendron infernense^[63]	Sp. nov	Valid	Manchester, Kvaček & Judd	Paleocene		United States ( Wyoming)	A species of Trochodendron. Announced in 2020; the final version of the article naming it was published in 2021.

Vitales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Yua jiangxiensis^[64]	Sp. nov	Valid	He & Wang	Miocene	Toupi Formation	China	A species of Yua. Announced in 2020; the final version of the article naming it was published in 2021.

Other angiosperms

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Aextoxicoxylon kawasianus^[65]	Sp. nov	Valid	Vera et al.	Late Cretaceous	Puntudo Chico Formation	Argentina	A fossil dicot wood
Atlantocarpus^[66]	Gen. et sp. nov	Valid	Friis, Crane & Pedersen	Early Cretaceous (Albian)	Potomac Group	Portugal United States ( Virginia)	An early flowering plant, possibly related to the group Austrobaileyales. Genus includes new species A. virginiensis.
Carpolithes gergoei^[13]	Sp. nov	Valid	Hably & Erdei in Hably	Miocene		Hungary	A fossil fruit of a flowering plant of uncertain phylogenetic placement.
Catanthus^[67]	Gen. et sp. nov	In press	Friis, Crane & Pedersen	Early Cretaceous		Portugal	An early flowering plant. Genus includes new species C. dolichostemon.
Cavilignum^[68]	Gen. et sp. nov	Valid	Siegert & Hermsen	Early Pliocene	Gray Fossil Site	United States ( Tennessee)	A flowering plant of uncertain phylogenetic placement, described on the basis of fossil endocarps. Genus includes new species C. pratchettii.
Chainandra^[69]	Gen. et sp. nov	Valid	Poinar & Chambers	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	Genus includes new species C. zeugostylus.
Cichoreacidites? flammulatus^[10]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin	Brazil	Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.
Crotonoidaepollenites echinatus^[10]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin	Brazil	Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.
Cyathitepala^[70]	Gen. et sp. nov	Valid	Poinar & Chambers	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	Genus includes new species C. papillosa.
Dasykothon^[71]	Gen. et sp. nov	Valid	Poinar & Chambers	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A flowering plant of uncertain phylogenetic placement, possibly a member of Laurales. Genus includes new species D. leptomiscus.
Dinganthus^[72]	Gen. et sp. nov	Valid	Liu et al.	Miocene	Dominican amber	Dominican Republic	A eudicot of uncertain phylogenetic placement. Genus includes new species D. pentamera.
Echistephanoporites annulatus^[10]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin	Brazil	Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.
Eofructus^[73]	Gen. et sp. nov	Valid	Han & Wang	Early Cretaceous	Yixian Formation	China	An infructescence including a central axis and five fruits resembling Liaoningfructus. Genus includes new species E. liutiaogouensis.
Foveotricolporites crassus^[10]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin	Brazil	Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.
Inaperturopollenites microechinatus^[10]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin	Brazil	Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.
Ladakhipollenites carmoi^[10]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin	Brazil	Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.
Lambertiflora^[66]	Gen. et sp. nov	Valid	Friis, Crane & Pedersen	Early Cretaceous (Albian)	Potomac Group	United States ( Virginia)	An early flowering plant, possibly related to the group Austrobaileyales. Genus includes new species L. elegans.
Malvacipolloides diversus^[10]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin	Brazil	Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.
Malvacipolloides echibaculatus^[10]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin	Brazil	Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.
Malvacipolloides romeroae^[10]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin	Brazil	Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.
Menatanthus^[74]	Gen. et sp. nov	Valid	Uhl, Paudayal & El Atfy in Uhl et al.	Paleocene (Thanetian)		France	A eudicot of uncertain phylogenetic placement. Genus includes new species M mosbruggeri. Announced in 2020; the final version of the article naming it was published in 2021.
Mugideiriflora^[66]	Gen. et sp. nov	Valid	Friis, Crane & Pedersen	Early Cretaceous (Aptian-early Albian)	Almargem Formation	Portugal	An early flowering plant, possibly related to the group Austrobaileyales. Genus includes new species M. portugallica.
Phantophlebia^[75]	Gen. et sp. nov	Valid	Poinar & Chambers	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A flowering plant of uncertain phylogenetic placement, possibly related to myrsinoid members of the family Primulaceae. Genus includes new species P. dicycla.
Platanites willigeri^[76]	Sp. nov	Valid	Halamski & Kvaček in Halamski et al.	Late Cretaceous (Santonian)	Czerna Formation	Poland	Trifoliolate platanoid leaves.
Psilaperiporites delicatus^[10]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin	Brazil	Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.
Psilaperiporites lunaris^[10]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin	Brazil	Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.
Ranunculacidites reticulatus^[10]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin	Brazil	Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.
Rasenganus^[77]	Gen. et sp. nov	Valid	Xing & Gu	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A possible epizoochorous fruit. Genus includes new species R. auricularus.
Retibrevitricolpites microreticulatus^[10]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin	Brazil	Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.
Retibrevitricolporites costaporus^[10]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin	Brazil	Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.
Retibrevitricolporites? toigoae^[10]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin	Brazil	Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.
Retipollenites solimoensis^[10]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin	Brazil	Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.
Retitriporites crassoreticulatus^[10]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin	Brazil	Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.
Rhoipites alfredii^[10]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin	Brazil	Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.
Singpuria^[78]	Gen. et sp. nov	Valid	Ramteke, Manchester & Nagrale in Ramteke et al.	Late Cretaceous (Maastrichtian)	Deccan Intertrappean Beds	India	A member of Pentapetalae of uncertain phylogenetic placement. Genus includes new species S. kapgatei.
Sinoherba^[79]	Gen. et sp. nov	Valid	Liu & Wang in Liu, Chen & Wang	Early Cretaceous (Barremian–Aptian)	Yixian Formation	China	An early monocot. Genus includes new species S. ningchengensis. Announced in 2020; the final version of the article naming it was published in 2021.
Varifructus^[80]	Gen. et sp. nov	In press	Liu et al.	Early Cretaceous	Yixian Formation	China	An early flowering plant. Genus includes new species V. lingyuanensis.
Wireroadia^[81]	Gen. et sp. et comb. nov	Valid	Zhang et al.	Late Cretaceous (Cenomanian to Santonian)		United States ( Alabama Massachusetts New Jersey New York)	A winged fruit of a eudicot of uncertain phylogenetic placement. Genus includes new species W. viccallii, as well as W. major (Hollick).

Pinales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Agathis ledongensis^[82]	Sp. nov	Valid	Oskolski et al.	Late Oligocene–early Miocene	Qiutangling Formation	China	A species of Agathis
Agathoxylon cozzoi^[83]	Sp. nov	Valid	Gnaedinger & Zavattieri	Late Triassic	Chihuido Formation	Argentina
Araucaria cuneoi^[84]	Sp. nov	Valid	Noll & Kunzmann	Middle Jurassic		Argentina	A species of Araucaria.
Araucaria famii^[85]	Sp. nov	Valid	Stockey & Rothwell	Late Cretaceous (Campanian)		Canada ( British Columbia)	A species of Araucaria.
Araucaria fildesensis^[86]	Sp. nov	Valid	Shi et al.	Eocene	Fossil Hill Formation	Antarctica (King George Island)	A species of Araucaria
Araucaria huncoensis^[87]	Sp. nov	Valid	Rossetto-Harris in Rossetto-Harris et al.	Early Eocene	Laguna del Hunco Formation	Argentina	A species of Araucaria.
Araucaria stockeyana^[84]	Sp. nov	Valid	Noll & Kunzmann	Middle Jurassic		Argentina	A species of Araucaria.
Araucarites pachacuteci^[88]	Sp. nov	In press	Martínez in Martínez et al.	Early Cretaceous (Berriasian–Valanginian)	Huancané Formation	Peru	A member of the family Araucariaceae
Brachyoxylon zhouii^[89]	Sp. nov	Valid	Jiang et al.	Early Cretaceous	Guantou Formation	China	A conifer wood. Announced in 2020; the final version of the article naming it was published in 2021.
Brachyphyllum sattlerae^[90]	Sp. nov	Valid	Batista et al.	Early Cretaceous (Aptian)	Crato Formation	Brazil	A member of the family Pinidae.
Callialastrobus^[91]	Gen. et sp. nov	In press	Kvaček & Mendes	Early Cretaceous (late Aptian–early Albian)	Lusitanian Basin	Portugal	A pollen cone of a member of Araucariaceae. Genus includes new species C. sousai.
Cedrus anatolica^[92]	Sp. nov	Valid	Akkemik	Early Miocene	Hançili Formation	Turkey	A species of Cedrus. Announced in 2020; the final version of the article naming it was published in 2021.
Circoporoxylon tibetense^[93]	Sp. nov	In press	Xia et al.	Middle Jurassic (Callovian)	Xiali Formation	China	Possibly a member of the family Podocarpaceae.
Comoxostrobus^[94]	Gen. et sp. nov	Valid	Stockey, Rothwell & Atkinson	Late Cretaceous (early Coniacian)		Canada ( British Columbia)	A member of the family Cupressaceae belonging to the subfamily Taiwanioideae. Genus includes new species C. rossii.
Cupressinoxylon klimovii^[95]	Nom. nov	Valid	Blokhina	Miocene		Russia	A member of the family Cupressaceae; a replacement name for Cupressinoxylon biotoides Blokhina (1989).
Cupressinoxylon llantenesense^[83]	Sp. nov	Valid	Gnaedinger & Zavattieri	Late Triassic	Llantenes Formation	Argentina
Cupressinoxylon manuelii^[96]	Sp. nov	In press	Ríos-Santos, Cevallos-Ferriz & Pujana	Late Cretaceous (Campanian-Maastrichtian)	Cabullona Group	Mexico
Ductoagathoxylon wangii^[97]	Sp. nov	In press	Gou & Feng in Gou et al.	Middle Jurassic	Xishanyao Formation	China	A conifer stem.
Friisia^[98]	Gen. et sp. nov	In press	Mendes & Kvaček	Early Cretaceous (late Aptian – early Albian)	Lusitanian Basin	Portugal	An ovuliferous cone of a member of the family Podocarpaceae. Genus includes new species F. lusitanica.
Juniperoxylon acarcaea^[99]	Sp. nov	In press	Akkemik	Early Miocene	Hançili Formation	Turkey	A member of the family Cupressaceae.
Lesbosoxylon kemaliyensis^[100]	Sp. nov	Valid	Akkemik & Mantzouka in Akkemik, Mantzouka & Kıran Yıldırım	Miocene	Divriği Formation	Turkey	A member of the family Pinaceae.
Marskea cuspidata^[101]	Sp. nov	Valid	Frolov & Mashchuk	Middle Jurassic	Prisayan Formation	Russia	A member of the family Taxaceae.
Mukawastrobus^[102]	Gen. et sp. nov	Valid	Stockey, Nishida & Rothwell	Late Cretaceous (late Campanian—early Maastrichtian)		Japan	A member of the family Cupressaceae belonging to the subfamily Taiwanioideae. Genus includes new species M. satoi.
Piceoxylon yumeniense^[103]	Sp. nov	Valid	Zhou, Peng, Deng, Zhang & Yang in Zhou et al.	Early Cretaceous	Xiagou Formation	China	Fossil wood of a member of the family Pinaceae. Announced in 2020; the final version of the article naming it was published in 2021.
Pinuxylon selmeierianum^[104]	Sp. nov	Valid	Dolezych & Reinhardt	Paleogene	Eureka Sound Group	Canada ( Nunavut)	A member of the family Pinaceae described on the basis of fossil wood
Protophyllocladoxylon chijinense^[103]	Sp. nov	Valid	Zhou, Peng, Deng, Zhang & Yang in Zhou et al.	Early Cretaceous	Xiagou Formation	China	Announced in 2020; the final version of the article naming it was published in 2021.
Tsuga asiatica^[105]	Sp. nov	Valid	Wu & Zhou in Wu et al.	Late Paleogene		China	A species of Tsuga. Announced in 2019; the final version of the article naming it was published in 2020.

Other seed plants

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Amyelon turpanense^[106]	Sp. nov	In press	Shi, Yu & Sun	Permian (Lopingian)	Wutonggou	China	A root of a member of Cordaitales
Androstrobus obovatus^[107]	Sp. nov	In press	Bodnar et al.	Late Triassic	Potrerillos Formation	Argentina	A member of Cycadales.
Araripestrobus^[108]	Gen. et sp. nov	Valid	Seyfullah, Roberts, Schmidt & Kunzmann in Seyfullah et al.	Early Cretaceous (Aptian-Albian)	Crato	Brazil	A seed plant belonging to the group Erdtmanithecales. Genus includes new species A. resinosus.
Archaeopetalanthus^[109]	Gen. et sp. nov	Valid	Naugolnykh	Carboniferous	Listvjanskaya	Russia	A member of Pinophyta belonging to the group Vojnovskyales. Genus includes new species A. progressus.
Battenispermum^[110]	Gen. et sp. nov	In press	Mendes, Pedersen & Friis	Early Cretaceous		Portugal	A seed plant belonging to the informal grouping Bennettitales-Erdtmanithecales-Gnetales. Genus includes new species B. hirsutum.
Carpolithus volantus^[111]	Sp. nov	Valid	Gómez et al.	Early Cretaceous (Aptian)	La Cantera Formation	Argentina	A fossil seed, possibly produced by a member of Gnetales.
Ductolobatopitys^[112]	Gen. et sp. nov	Valid	Conceição & Crisafulli in Conceição et al.	Permian (Cisuralian)	Pedra de Fogo Formation	Brazil	A gymnosperm described on the basis of fossil wood. Genus includes new species D. mussae.
Filigigantopteris^[113]	Gen. et 2 sp. nov	In press	Zhou et al.	Late Permian	Nayixiong Formation	China	A gigantopterid. Genus includes new species F. asymmetrica and F. dahaia.
Ginkgo pediculata^[114]	Sp. nov	In press	Deng, Yang & Zhou	Early Cretaceous		China	A species of Ginkgo.
Jianchangia^[115]	Gen. et sp. nov	Valid	Yang, Wang & Ferguson	Early Cretaceous (Aptian)	Jiufotang Formation	China	A member of Ephedraceae. Genus includes new species J. verticillata.
Johniphyllum^[116]	Gen. et sp. nov	Valid	Looy & Duijnstee	Permian (Guadalupian)		United States ( Texas)	A member of Voltziales. Genus includes new species J. multinerve.
Jordaniopteris^[117]	Gen. et comb. nov	Valid	Anderson in Anderson et al.	Permian (possibly Lopingian)	Um Irna Formation	Jordan	A seed fern. A new genus for "Dicroidium" irnensis Abu Hamad et al. (2008); genus also includes "Dicroidium" jordanensis Abu Hamad et al. (2008), "Dicroidium" robustum Kerp & Vörding (2008) and "Dicroidium" bandelii Abu Hamad et al. (2017).
Kaokoxylon brasiliensis^[112]	Sp. nov	Valid	Conceição, Neregato & Iannuzzi in Conceição et al.	Permian (Cisuralian)	Pedra de Fogo Formation	Brazil	A conifer described on the basis of fossil wood.
Nilssonia mirovanae^[118]	Sp. nov	Valid	Čepičková & Kvaček	Late Cretaceous (Cenomanian)	Peruc-Korycany Formation	Czech Republic	A cycad.
Novaiorquepitys^[119]	Gen. et sp. nov	In press	Conceição & Crisafulli in Conceição et al.	Permian (Cisuralian)	Pedra de Fogo Formation	Brazil	A gymnosperm stem. Genus includes new species N. maranhensis Conceição, Neregato & Iannuzzi.
Ovalocarpus butmanii^[120]	Sp. nov	Valid	Naugolnykh & Linkevich	Permian (Artinskian)		Russia ( Sverdlovsk Oblast)	A member of Ginkgoales belonging to the family Cheirocladaceae.
Palaeocupressinoxylon^[121]	Gen. et sp. nov	In press	Wan, Yang & Wang	Late Permian	Turpan–Hami Basin	China	A silicified gymnospermous fossil wood. Genus includes new species P. uniseriale.
Phoenicopsis anadyrensis^[122]	Sp. nov	Valid	Nosova in Zolina et al.	Late Cretaceous–Paleocene (Maastrichtian–Danian)	Rarytkin	Russia ( Chukotka Autonomous Okrug)	A member of Czekanowskiales
Pseudovoltzia sapflorensis^[116]	Sp. nov	Valid	Looy & Duijnstee	Permian (Guadalupian)		United States ( Texas)	A member of Voltziales
Pterostoma neehoffii^[123]	Sp. nov	Valid	Conran et al.	Middle Miocene		New Zealand	A cycad
Pteruchus frenguellii^[124]	Sp. nov	In press	Blomenkemper et al.	Late Permian	Umm Irna Formation	Jordan	A pollen organ of a seed fern
Pteruchus lepidus^[124]	Sp. nov	In press	Blomenkemper et al.	Late Permian	Umm Irna Formation	Jordan	A pollen organ of a seed fern
Samaropsis jinchangensis^[125]	Sp. nov	Valid	Hua & Sun in Hua et al.	Early Permian		China	A seed fossil. Announced in 2019; the final version of the article naming was published in 2020.
Umaltolepis involuta^[126]	Sp. nov	In press	Nosova	Middle Jurassic	Angren Formation	Uzbekistan
Umaltolepis sogdianica^[126]	Sp. nov	In press	Nosova	Middle Jurassic	Angren Formation	Uzbekistan
Umkomasia aequatorialis^[124]	Sp. nov	In press	Blomenkemper et al.	Late Permian	Umm Irna Formation	Jordan	A cupulate structure of a seed fern
Wantus^[116]	Gen. et sp. nov	Valid	Looy & Duijnstee	Permian (Guadalupian)		United States ( Texas)	A member of Voltziales. Genus includes new species W. acaulis.
Wudaeophyton^[127]	Gen. et sp. nov	Valid	Pšenička et al.	Early Permian	Taiyuan Formation	China	A small vine, most similar to pteridosperms from the group Callistophytales. Genus includes new species W. wangii.
Yangopteris^[128]	Gen. et comb. nov	In press	Zhou et al.	Permian (Asselian)		China	A seed fern; a new genus for "Alethopteris" ascendens Halle.
Yvyrapitys^[119]	Gen. et sp. nov	In press	Conceição & Crisafulli in Conceição et al.	Permian (Cisuralian)	Pedra de Fogo Formation	Brazil	A gymnosperm stem. Genus includes new species Y. novaiorquensis Conceição, Neregato & Iannuzzi.

Other plants

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Annularia paisii^[129]	Sp. nov	Valid	Correia et al.	Carboniferous (Gzhelian)		Portugal
Blasiites huolinhensis^[130]	Sp. nov	In press	Li et al.	Early Cretaceous	Huolinhe Formation	China	A liverwort belonging to the family Blasiaceae.
Botryopteris multifolia^[131]	Sp. nov	In press	He et al.	Permian (Lopingian)	Junlian Formation	China	A fern
Birisia mandshurica^[132]	Sp. nov	Valid	Golovneva, Grabovskiy & Zolina	Early Cretaceous (Albian)	Frentsevka Formation	Russia ( Primorsky Krai)	A fern belonging to the family Dicksoniaceae.
Calamites cambrensis^[133]	Sp. nov	Valid	Thomas	Carboniferous (Pennsylvanian)		United Kingdom
Catenuporella^[134]	Gen. et sp. nov	Valid	Zhang et al.	Late Ordovician	Ordos Basin	China	A green alga belonging to the group Dasycladales. Genus includes new species C. gigantia.
Charaxis spicatus^[135]	emend. nov.	Valid	Pérez-Cano, Bover-Arnal et Martín-Closas in Pérez-Cano et al.	Barremian	Maestrat Basin	Spain	Thallus of Echinochara lazarii. Both taxa has been firstly found anatomically attached
Chlamydomonas hanublikanus^[136]	Sp. nov		Vršanská & Hinkelman	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A species of Chlamydomonas
Circinites^[47]	Gen. et sp. nov	In press	Robledo & Anzótegui in Robledo et al.	Miocene-Pliocene		Argentina	A fern belonging to the family Pteridaceae. Genus includes new species C. pteridoides.
Clavatisporites cenomaniana^[137]	Sp. nov	Valid	Santamarina in Santamarina et al.	Late Cretaceous (Cenomanian)	Mata Amarilla Formation	Argentina	Spores of a member of Filicopsida of uncertain phylogenetic placement. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.
Clavator calcitrapus var. jiangluoensis^[135]	comb. nov	Valid	Pérez-Cano, Bover-Arnal et Martín-Closas in Pérez-Cano et al.	Barremian	Maestrat Basin	Spain	Clavatoracean species.
Collarecodium? nezpercae^[138]	Sp. nov	Valid	Bucur & Rigaud in Bucur et al.	Late Triassic (Norian)		United States ( Idaho)	A green alga belonging to the group Bryopsidales and possibly to the family Udoteaceae.
Collarisporites minor^[137]	Sp. nov	Valid	Santamarina in Santamarina et al.	Late Cretaceous (Cenomanian)	Mata Amarilla Formation	Argentina	Spores of a member of Filicopsida of uncertain phylogenetic placement. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.
Coniopteris sandaolingensis^[139]	Sp. nov	Valid	Yuan & Sun in Yuan et al.	Middle Jurassic	Xishanyao Formation	China
Dimicheleodendron^[140]	Gen. et comb. nov	Valid	Thomas & Cleal	Carboniferous		United Kingdom	A lycophyte; a new genus for "Lepidodendron" hickii.
Drynaria diplosticha^[141]	Sp. nov	Valid	Yu & Xie in Yu et al.	Late Miocene	Bangmai Formation	China	A fern belonging to the family Polypodiaceae.
Echinochara lazarii^[135]	comb. nov	Valid	Pérez-Cano, Bover-Arnal et Martín-Closas in Pérez-Cano et al.	Barremian	Maestrat Basin	Spain	A member of Clavatoraceans.
Equicalastrobus pusillus^[142]	Sp. nov	Valid	Zhang & Yan in Zhang et al.	Late Triassic	Baojishan Basin	China	A member of Equisetales. Announced in 2020; the final version of the article naming was published in 2021.
Equisetum yenbaiense^[143]	Sp. nov	Valid	Aung et al.	Late Miocene		Vietnam	A species of Equisetum
Equisetum yongpingense^[143]	Sp. nov	Valid	Aung et al.	Late Pliocene	Sanying Formation	Vietnam	A species of Equisetum
Filippoporella^[144]	Gen. et sp. nov	In press	Sokač & Grgasović	Early Paleocene		Croatia	A green alga belonging to the group Dasycladales. Genus includes new species F. barattoloi.
Frullania partita^[145]	Sp. nov	Valid	Li et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A liverwort, a species of Frullania
Frullania vanae^[146]	Sp. nov	Valid	Mamontov et al.	Eocene	Rovno amber	Ukraine	A liverwort, a species of Frullania
Gippslandites^[147]	Gen. et sp. nov	Valid	McSweeney, Shimeta & Buckeridge	Late Silurian–early Devonian		Australia	A member of Zosterophyllaceae. Genus includes new species G. minutus.
Gmujij^[148]	Gen. et sp. nov	Valid	Pfeiler & Tomescu	Devonian (Emsian)	Battery Point	Canada ( Quebec)	An early euphyllophyte. Genus includes new species G. tetraxylopteroides. Announced in 2020; the final version of the article naming it was published in 2021.
Griphoporella minuta^[138]	Sp. nov	Valid	Bucur & Peybernes in Bucur et al.	Late Triassic		Japan	A green alga belonging to the group Dasycladales and the family Triploporellaceae.
Hansopteris^[149]	Gen. et sp. nov	In press	Zhou et al.	Permian (Asselian)		China	An anachoropterid fern. Genus includes new species H. uncinatus.
Holosporella magna^[138]	Sp. nov	Valid	Bucur & Fucelli in Bucur et al.	Late Triassic (Norian)		United States ( Nevada)	A green alga belonging to the group Dasycladales and the family Triploporellaceae.
Holosporella? rossanae^[138]	Sp. nov	Valid	Bucur & Del Piero in Bucur et al.	Late Triassic (Norian)		Canada ( Yukon)	A green alga belonging to the group Dasycladales and the family Triploporellaceae.
Inocladus^[150]	Gen. et comb. nov	Valid	LoDuca et al.	Silurian		United States	An alga related to the group Bryopsidales. Genus includes "Buthotrephis" divaricata White (1901), "B." newlini White (1901), "B." lesquereuxi Grote & Pitt (1876) and "Chondrites" verus Ruedemann (1925). Announced in 2020; the final version of the article naming it was published in 2021.
Intermurella ordosensis^[134]	Sp. nov	Valid	Zhang et al.	Late Ordovician	Ordos Basin	China	A green alga belonging to the group Dasycladales.
Jurafructus^[151]	Gen. et sp. nov	Valid	Chen et al.	Middle−Late Jurassic	Jiulongshan Formation	China	A plant of uncertain phylogenetic placement, possibly a flowering plant described on the basis of a probable fossil drupe. Genus includes new species J. daohugouensis.
Keraphyton^[152]	Gen. et sp. nov	Valid	Champreux, Meyer-Berthaud & Decombeix	Devonian (Famennian)	Mandowa Mudstone Formation	Australia	A member of Iridopteridales of uncertain phylogenetic placement. Genus includes new species K. mawsoniae.
Khasurtya^[153]	Gen. et sp. nov	Valid	Mamontov in Kopylov et al.	Early Cretaceous		Russia	A liverwort belonging to the group Marchantiidae. Genus includes new species K. ginkgoides.
Lepidodendron demkinae^[154]	Sp. nov	Valid	Mosseichik	Carboniferous (Viséan)		Russia
Lobatannularia linjiaensis^[155]	Sp. nov	Valid	Xu et al.	Middle Triassic	Linjia Formation	China	A member of Equisetales.
Lygodium sanshuiense^[156]	Sp. nov	Valid	Naugolnykh et al.	Paleocene	Buxin Formation	China	A species of Lygodium. Announced in 2019; the final version of the article naming it was published in 2020.
Metzgeriites kujiensis^[157]	Sp. nov	Valid	Katagiri in Katagiri & Shinden	Late Cretaceous (Santonian)	Tamagawa Formation	Japan	A liverwort.
Munieria martinclosasi^[135]	comb. nov.	Valid	Pérez-Cano, Bover-Arnal et Martín-Closas in Pérez-Cano et al.	Barremian	Lebanon	Lebanon	Clavatoracean thallus. Formerly known as Charaxis martinclosasi
Neoarthropitys^[158]	Gen. et sp. nov	In press	Gnaedinger et al.	Middle Triassic	Quebrada de los Fósiles	Argentina	A member of Equisetales. Genus includes new species N. gondwanaensis.
Osmundacaulis asiatica^[159]	Sp. nov	Valid	Cheng et al.	Cretaceous		China	A member of the family Osmundaceae
Osmundacaulis sinica^[159]	Sp. nov	Valid	Cheng et al.	Cretaceous		China	A member of the family Osmundaceae
Ovoidites circumplicatus^[160]	Sp. nov	Valid	Zavattieri, Gutiérrez & Monti	Middle Triassic	Quebrada de los Fósiles Formation	Argentina	A green alga belonging to the group Zygnematales.
Ovoidites tripartitus^[160]	Sp. nov	Valid	Zavattieri, Gutiérrez & Monti	Middle Triassic	Quebrada de los Fósiles Formation	Argentina	A green alga belonging to the group Zygnematales.
Palaeostachya guanglongii^[161]	Sp. nov	In press	Liu et al.	Permian (Asselian)	Taiyuan Formation	China	A member of the family Calamitaceae.
Parazosterophyllum^[147]	Gen. et sp. nov	Valid	McSweeney, Shimeta & Buckeridge	Late Silurian–early Devonian		Australia	A member of Zosterophyllaceae. Genus includes new species P. timsiae.
Patruliuspora oregonica^[138]	Sp. nov	Valid	Bucur & Rigaud in Bucur et al.	Late Triassic (Norian)		United States ( Oregon)	A green alga belonging to the group Dasycladales and the family Polyphysaceae.
Patruliuspora pacifica^[138]	Sp. nov	Valid	Bucur, Del Piero & Peyrotty in Bucur et al.	Late Triassic (Norian)		Canada ( Yukon)	A green alga belonging to the group Dasycladales and the family Polyphysaceae.
Pellites hamiensis^[162]	Sp. nov	Valid	Li et al.	Middle Jurassic	Xishanyao	China	A liverwort belonging to the family Pelliaceae.
Plenasium (Aurealcaulis) elegans^[163]	Sp. nov	In press	Hiller et al.	Eocene	Na Duong Formation	Vietnam	A member of Osmundaceae
Pleuromeia shaolinii^[164]	Sp. nov	Valid	Zhang & Wang in Zhang et al.	Middle Triassic	Linjia	China
Polycingulatisporites multiverrucata^[137]	Sp. nov	Valid	Santamarina in Santamarina et al.	Late Cretaceous (Cenomanian)	Mata Amarilla	Argentina	Spores of a member of Bryophyta of uncertain phylogenetic placement, possibly of sphagnaceous affinity. Announced in 2019; the final version of the article naming it was published in 2020.
Polypodiisporites minutiverrucatus^[10]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin	Brazil	Pteridophyte spore. Announced in 2020; the final version of the article naming it was published in 2021.
Polysporia baetica^[165]	Sp. nov	In press	Álvarez-Vázquez, Bek & Drábková	Carboniferous (Pennsylvanian)	Peñarroya-Belmez-Espiel Coalfield	Spain	A member of Isoetales
Polystichum pacltovae^[166]	Sp. nov	Valid	Kvaček in Kvaček & Teodoridis	Oligocene		Czech Republic	A fern, a species of Polystichum
Proodontosoria^[167]	Gen. et sp. nov	Valid	Li et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A fern belonging to the family Lindsaeaceae. Genus includes new species P. myanmarensis.
Proterocladus antiquus^[168]	Sp. nov	Valid	Tang et al.	Mesoproterozoic circa 1 Ga	Nanfen	China	An early siphonocladalean chlorophyte
Psilatriletes cozzuolii^[10]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin	Brazil	Pteridophyte spore. Announced in 2020; the final version of the article naming it was published in 2021.
Psilochara monevaensis^[169]	Sp. nov	Valid	Sanjuan & Soulié-Märsche	Middle Miocene		Spain	A charophyte.
Qianshouia^[170]	Gen. et sp. nov	Valid	Huang et al.	Late Devonian	Wutong Formation	China	A plant of uncertain phylogenetic placement, possibly a lycopsid or a sphenopsid. Genus includes new species Q. mira.
Scolecopteris minuta^[171]	Sp. nov	In press	Wan et al.	Early Permian	Taiyuan Formation	China	A fern belonging to the group Marattiales.
Sigillaria pfefferkornii^[172]	Sp. nov	In press	D'Antonio, Boyce & Wang	Permian (Asselian)		China
Sigillaria wudensis^[172]	Sp. nov	In press	D'Antonio, Boyce & Wang	Permian (Asselian)		China
Sphaerochara miocenica^[169]	Sp. nov	Valid	Sanjuan & Soulié-Märsche	Miocene		Lebanon Spain	A charophyte.
Thyrsopteris cretacea^[173]	Sp. nov	Valid	Li et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A species of Thyrsopteris
Tomiostrobus sinensis^[174]	Sp. nov	Valid	Feng in Feng et al.	Early Triassic (Induan)	Kayitou Formation	China	A member of the family Isoetaceae.
Tumidopteris astra^[175]	Sp. nov	Valid	Naugolnykh	Permian (Roadian)	Pechora coal basin	Russia	A fern belonging to the family Gleicheniaceae.
Ufadendron elongatum^[176]	Sp. nov	Valid	Tang et al.	Late Permian	Linxi	China	A lycopsid belonging to the family Tomiodendraceae
Uzhurodendron^[177]	Gen. et sp. nov	Valid	Mosseichik & Filimonov	Carboniferous (Tournaisian)	Bystrianskaya	Russia ( Krasnoyarsk Krai)	A member of Lycopodiopsida. Genus includes new species U. asiaticum.
Zeilleria fosteri^[178]	Sp. nov	Valid	Thomas et al.	Carboniferous (Bashkirian)		United Kingdom	A fern

General research

A study on the evolutionary history of green plants is published by Nie et al. (2020).^[179]
Description of new fossil material of Yurtusia uniformis from the Cambrian Yanjiahe Formation (China) and a study on the phylogenetic relationships and possible life cycle of this organism is published by Shang et al. (2020), who consider Y. uniformis to be a likely green microalga.^[180]
A study on the phylogenetic relationships of extant and fossil complex thalloid liverworts (Marchantiidae) is published by Flores et al. (2020).^[181]
Evidence of development of dichotomous roots in euphyllophytes that were extant during the Devonian and Carboniferous periods is presented by Hetherington, Berry & Dolan (2020), who interpret their findings as indicating that dichotomous root branching evolved in both lycophytes and euphyllophytes.^[182]
An early land plant producing multiple spore size classes is described from the Lower Devonian Campbellton Formation (Canada) by Bonacorsi et al. (2020).^[183]
A study on the impact of the appearance and evolution of herbivorous tetrapods on the evolution of land plants from the Carboniferous to the Early Triassic is published by Brocklehurst, Kammerer & Benson (2020).^[184]
A study on the production of periderm in Late Paleozoic arborescent lycopsids is published by D'Antonio & Boyce (2020), who argue that these lycopsids did not grow from sporelings into large trees through the production of a periderm cylinder, because physiological limitations would have prohibited the production of thick periderm.^[185]
A study on the architecture and development of the Carboniferous arborescent lycopsid Paralycopodites is published by DiMichele & Bateman (2020).^[186]
New information on the anatomy of Weichselia reticulata is presented by Blanco-Moreno, Decombeix & Prestianni (2020).^[187]
A study on the phylogenetic placement of the extinct fern genus Coniopteris is published by Li et al. (2020).^[188]
New information on the morphology of Paleoazolla patagonica is presented by Benedetti et al. (2020), who evaluate the implications of this taxon for the knowledge of the evolution of water ferns.^[189]
A study aiming to determine which ferns were most likely to be the producers of Cyathidites spores from earliest Paleocene plant localities across western North America, and were most likely to be among the first plants in western North America to thrive in the immediate aftermath of the Cretaceous–Paleogene extinction event, is published by Berry (2020).^[190]
A study on the morphology and development of Genomosperma, and on its implications for the knowledge of the evolutionary origins of seed development, is published by Meade, Plackett & Hilton (2020).^[191]
A pollen organ resembling seed fern pollen organs Dictyothalamus and Melissiotheca is described from the Lopingian Umm Irna Formation (Jordan) by Zavialova et al. (2020), who interpret this finding as evidence of persistence of lyginopterid seed ferns until the late Permian.^[192]
Evidence of increasing atmospheric CO₂ concentration at the onset of the end-Triassic extinction event, based on data from fossil leaves of the seed fern Lepidopteris ottonis from southern Sweden, is presented by Slodownik, Vajda & Steinthorsdottir (2020), who confirm L. ottonis as a valid proxy for pCO₂ reconstructions.^[193]
A study on the anatomy of the seed cone scales of Krassilovia mongolica is published by Herrera et al. (2020), who argue that K. mongolica and Podozamites harrisii are the seed cones and leaves of the same extinct plant, and name a new family Krassiloviaceae within the order Voltziales.^[194]
A study on the microscopic wood anatomy of a fossil tree trunk of Agathoxylon arizonicum with the characteristic external features of a fire scar from the Upper Triassic Chinle Formation (Petrified Forest National Park, Arizona, United States) is published by Byers et al. (2020), who evaluate the implications of this specimen for the knowledge of the evolution of fire-adapted plant traits.^[195]
A putative bamboo "Chusquea" oxyphylla from the early Eocene Laguna del Hunco biota (Argentina) is reinterpreted as a conifer by Wilf (2020), who transfers this species to the genus Retrophyllum.^[196]
A study on evolutionary history of conifers as indicated by fossil and molecular data, aiming to determine whether the rise of angiosperms drove the decline of conifers and other gymnosperms, is published by Condamine et al. (2020).^[197]
Presence of secretory tissues is reported in extinct flowers from the Cretaceous amber from Myanmar and Cenozoic Dominican amber (including specimens preserved while in the process of emitting compounds) by Poinar & Poinar (2020).^[198]
Fossil pollen of flowering plants is reported from the Aptian and Albian of Australia by Korasidis & Wagstaff (2020), who evaluate the implications of their findings for the knowledge of the timing of the appearance and diversification of the flowering plants in the high-latitude southern basins of Australia.^[199]
A study on the morphology of palm and palm-like pollen from the Eocene Yaw Formation (Myanmar), and on the implications of these fossils for the knowledge of distribution and diversity of Eocene palms across the globe, is published by Huang et al. (2020).^[200]
Fossils fruits of Illigera eocenica, representing the second fossil occurrence of Illigera worldwide and the first in Asia, are described from the Eocene Niubao Formation (central Tibetan Plateau) by Wang et al. (2020), who evaluate the implications of this finding for the knowledge of the climate in the central Tibetan Plateau during the early middle Eocene, and for the knowledge of the floristic links between Asia and North America during the Paleogene.^[201]
A study on the morphology and phylogenetic relationships of Montsechia vidalii is published by Gomez et al. (2020).^[202]
Eocene leaves of members of the family Urticaceae with stinging trichomes are described from the Okanogan Highlands (British Columbia, Canada) by DeVore et al. (2020).^[203]
A revision of the fossil record of the family Nothofagaceae from South America is published by Pujana et al. (2020).^[204]
A study on the extinction of plants from south polar terrestrial ecosystems during the Permian–Triassic extinction event and on their recovery after this extinction event, based on data from the Sydney Basin (Australia), is published by Mays et al. (2020).^[205]
A study on the impact of ecological disturbances around the Permian–Triassic boundary (from the Wuchiapingian to Ladinian) on land plant communities is published by Nowak, Vérard & Kustatscher (2020).^[206]
A study on the age of the Paleogene Kanaka Creek fossil flora (Huntingdon Formation; British Columbia, Canada) and on its implications for reconstructions of the contemporaneous paleoclimate and paleoenvironment is published by Mathewes, Greenwood & Love (2020).^[207]
Evidence from Eocene plant fossils from the Bangong-Nujiang suture indicating that the Tibetan Plateau area hosted a diverse subtropical ecosystem approximately 47 million years ago and that this area was both low and humid at the time is presented by Su et al. (2020), who also report that the composition of this flora is similar to Early-Middle Eocene floras in both North America and Europe, but shows little affinity to Eocene floras from the Indian Plate.^[208]
A study aiming to estimate leaf dry mass per area in fossil plants from 22 western North American sites spanning the Eocene–Oligocene transition is published online by Butrim & Royer (2020), who evaluate the implications of their findings for the knowledge of the impact of the environmental changes occurring during the Eocene–Oligocene transition on leaf-economic strategies of plants.^[209]
A study on the Neogene paleobotanical record and climate in the northernmost part of the Central Andean Plateau, based on data from the Descanso Formation (Peru), is published by Martínez et al. (2020), who report the earliest evidence of a puna-like ecosystem in the Pliocene and a montane ecosystem without modern analogs in the Miocene.^[210]
Fossil fruits (mericarps) of the neoendemic Apiaceae Melanoselinum (≡ Daucus) decipiens were reported from the lacustrine and fluvial sediments of Porto da Cruz, Madeira, dated 1.3 Ma, by Góis-Marques et al. 2020.^[211] This paper not only reports the oldest Daucus s.l. fossil known to date but also the first fossil evidence of a plant with insular woodiness (see Island gigantism).
The leaf fossil Mesodescolea plicata from the Early Cretaceous of Patagonia, first interpreted as a cycad with affinities with extant Stangeria, is reinterpreted as an angiosperm leaf with affinities with Austrobaileyales or Chloranthales by Coiro et al. 2020,^[212] with implications for the evolution of leaf shape in the early radiation of the angiosperms.
A study on the phylogenetic relationships of 10 Cretaceous flower taxa (Chloranthistemon endressii, Dakotanthus cordiformis, Kajanthus lusitanicus, Mauldinia mirabilis, Microvictoria svitkoana, Paleoclusia chevalieri, Paradinandra suecica, Spanomera mauldiniensis, Tylerianthus crossmanensis and Virginianthus calycanthoides) is published by Schönenberger et al. (2020).^[213]

References

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^ Ruth A. Stockey; Harufumi Nishida; Gar W. Rothwell (2020). "Evolutionary diversification of taiwanioid conifers: evidence from a new Upper Cretaceous seed cone from Hokkaido, Japan". Journal of Plant Research. 133 (5): 681–692. Bibcode:2020JPlR..133..681S. doi:10.1007/s10265-020-01214-y. PMC 7429551. PMID 32686035.
^ ^a ^b Ling-Qi Zhou; Cuo Peng; Peng Deng; Xiao-Qin Zhang; Guo-Lin Yang; Wen-Xiu Ren; Jun Wang; Xiao-Qiang Li; Shi-Bo Tuo; Bing Guo (2020). "New records of Early Cretaceous petrified wood in Yumen, northwestern Gansu Province, China and their palaeoclimatic implications". Palaeoworld. 30 (3): 503–514. doi:10.1016/j.palwor.2020.08.002.
^ Martina Dolezych; Lutz Reinhardt (2020). "First evidence for the conifer Pinus, as Pinuxylon selmeierianum sp. nov., during the Paleogene on Wootton Peninsula, northern Ellesmere Island, Nunavut, Canada". Canadian Journal of Earth Sciences. 57 (1): 25–39. Bibcode:2020CaJES..57...25D. doi:10.1139/cjes-2018-0163.
^ Meng-Xiao Wu; Jian Huang; Tao Su; Qin Leng; Zhe-Kun Zhou (2020). "Tsuga seed cones from the late Paleogene of southwestern China and their biogeographical and paleoenvironmental implications". Palaeoworld. 29 (3): 617–628. doi:10.1016/j.palwor.2019.07.005. S2CID 199885815.
^ Xiao Shi; Jianxin Yu; Yuewu Sun (2020). "Tyloses in the Lopingian cordaitalean root from Xinjiang, Northwest China". Review of Palaeobotany and Palynology. 273: Article 104134. Bibcode:2020RPaPa.27304134S. doi:10.1016/j.revpalbo.2019.104134. S2CID 210278581.
^ Josefina Bodnar; Eduardo M. Morel; Eliana P. Coturel; Daniel G. Ganuza (2020). "New plant fossil records and biostratigraphic analysis from the Uspallata Group (Late Triassic) at Cacheuta Hill, Cuyo Basin, west-central Argentina". Geobios. 60: 3–27. Bibcode:2020Geobi..60....3B. doi:10.1016/j.geobios.2020.04.002. S2CID 219762016.
^ Leyla J. Seyfullah; Emily A. Roberts; Alexander R. Schmidt; Eugenio Ragazzi; Ken B. Anderson; Daniel Rodrigues do Nascimento Jr.; Wellington Ferreira da Silva Filho; Lutz Kunzmann (2020). "Revealing the diversity of amber source plants from the Early Cretaceous Crato Formation, Brazil". BMC Evolutionary Biology. 20 (1): 107. Bibcode:2020BMCEE..20..107S. doi:10.1186/s12862-020-01651-2. PMC 7439571. PMID 32819273. S2CID 221183260.
^ Serge V. Naugolnykh (2020). "Archaeopetalanthus progressus gen. et sp. nov. – a new representative of the vojnovskyopsid gymnosperms from the Carboniferous of Siberia (Russia)". Wulfenia. 27: 97–113.
^ Mário Miguel Mendes; Kaj Raunsgaard Pedersen; Else Marie Friis (2020). "Battenispermum hirsutum gen. et sp. nov., a new Early Cretaceous seed from Portugal with chlamydospermous organisation". Cretaceous Research. 109: Article 104376. Bibcode:2020CrRes.10904376M. doi:10.1016/j.cretres.2020.104376. S2CID 213898064.
^ María A. Gómez; Gabriela G. Puebla; Mercedes B. Prámparo; Andrea B. Arcucci (2020). "Fossil seeds from the La Cantera Formation, Early Cretaceous, San Luis Province, Argentina". Acta Palaeobotanica. 60 (1): 181–198. doi:10.35535/acpa-2020-0008. hdl:11336/145016.
^ ^a ^b Domingas Maria da Conceição; Luiz Saturnino de Andrade; Rodrigo Neregato; Roberto Iannuzzi; Alexandra Crisafulli; Juan Carlos Cisneros (2020). "New petrified gymnosperms from the Permian of Maranhão (Pedra de Fogo Formation), Brazil: Ductolobatopitys nov. gen. and Kaokoxylon". Geobios. 60: 47–59. Bibcode:2020Geobi..60...47C. doi:10.1016/j.geobios.2020.04.003. S2CID 219737764.
^ Wei-Ming Zhou; Bi-Yun Chen; Wei Sun; Xue-Zhi He; Jason Hilton; Jun Wang (2020). "A new gigantopterid genus from the late Permian of the Daha Coalfield, Tibetan Plateau and its implication on plant-insect interactions". Historical Biology: An International Journal of Paleobiology. 33 (12): 3228–3240. doi:10.1080/08912963.2020.1860033. S2CID 234390424.
^ Sheng-Hui Deng; Xiao-Ju Yang; Zhi-Yan Zhou (2020). "A new Ginkgo from the Lower Cretaceous of Liaoning, Northeast China and its evolutionary implications". Review of Palaeobotany and Palynology. 283: Article 104315. Bibcode:2020RPaPa.28304315D. doi:10.1016/j.revpalbo.2020.104315. S2CID 225031049.
^ Yong Yang; Yingwei Wang; David Kay Ferguson (2020). "A new macrofossil ephedroid plant with unusual bract morphology from the Lower Cretaceous Jiufotang Formation of northeastern China". BMC Evolutionary Biology. 20 (1): Article number 19. Bibcode:2020BMCEE..20...19Y. doi:10.1186/s12862-019-1569-y. PMC 7001366. PMID 32019502.
^ ^a ^b ^c Cindy V. Looy; Ivo A. P. Duijnstee (2020). "Voltzian conifers of the South Ash Pasture flora (Guadalupian, Texas): Johniphyllum multinerve gen. et sp. nov., Pseudovoltzia sapflorensis sp. nov., and Wantus acaulis gen. et sp. nov". International Journal of Plant Sciences. 181 (3): 363–385. doi:10.1086/706853. S2CID 208592093.
^ Heidi M. Anderson; Maria Barbacka; Marion K. Bamford; W. B. Keith Holmes; John M. Anderson (2020). "Dicroidium (foliage) and affiliated wood Part 3 of a reassessment of Gondwana Triassic plant genera and a reclassification of some previously attributed". Alcheringa: An Australasian Journal of Palaeontology. 44 (1): 64–92. Bibcode:2020Alch...44...64A. doi:10.1080/03115518.2019.1622779. S2CID 199109037.
^ Jana Čepičková; Jiří Kvaček (2020). "Two cycads Nilssonia mirovanae sp. nov. and Pseudoctenis babinensis J.Kvaček from the Cenomanian of the Bohemian Cretaceous Basin (the Czech Republic) as indicators of water stress in the palaeoenvironment". Fossil Imprint. 76 (2): 315–324. doi:10.37520/fi.2020.025.
^ ^a ^b Domingas Maria da Conceição; Alexandra Crisafulli; Roberto Iannuzzi; Rodrigo Neregato; Juan Carlos Cisneros; Luiz Saturnino de Andrade (2020). "New petrified gymnosperms from the Permian of Maranhão (Pedra de Fogo Formation), Brazil: Novaiorquepitys and Yvyrapitys". Review of Palaeobotany and Palynology. 276: Article 104177. Bibcode:2020RPaPa.27604177D. doi:10.1016/j.revpalbo.2020.104177. S2CID 212954578.
^ Sergey V. Naugolnykh; Valeryi V. Linkevich (2020). "Artinskian flora (Lower Permian) of the stratotypic area (the Middle Cis-Urals)". Socialno-ecologicheskie Technologii. 10 (2): 133–150. doi:10.31862/2500-2961-2020-10-2-133-150.
^ Ming-Li Wan; Wan Yang; Jun Wang (2020). "Palaeocupressinoxylon uniseriale n. gen. n. sp., a gymnospermous wood from the upper Permian of Central Taodonggou, southern Bogda Mountains, northwestern China". Palaeoworld. 29 (1): 117–125. doi:10.1016/j.palwor.2019.06.002.
^ Anastasia Zolina; Lina Golovneva; Natalya Nosova; Alexander Grabovskiy (2020). "A new species of Phoenicopsis (Leptostrobales) from the Maastrichtian–Danian of Chukotka, Russia". Geobios. 63: 67–75. Bibcode:2020Geobi..63...67Z. doi:10.1016/j.geobios.2020.09.002. S2CID 228928283.
^ John G. Conran; Jennifer M. Bannister; Uwe Kaulfuss; Daphne E. Lee (2020). "Pterostoma neehoffii (cf. Zamiaceae): a new species of extinct cycad from the middle Miocene of New Zealand and an overview of fossil New Zealand cycads". New Zealand Journal of Botany. 58 (1): 30–47. Bibcode:2020NZJB...58...30C. doi:10.1080/0028825X.2019.1653939. S2CID 202847120.
^ ^a ^b ^c Patrick Blomenkemper; Hans Kerp; Abdalla Abu Hamad; Benjamin Bomfleur (2020). "Contributions towards whole-plant reconstructions of Dicroidium plants (Umkomasiaceae) from the Permian of Jordan". Review of Palaeobotany and Palynology. 278: Article 104210. Bibcode:2020RPaPa.27804210B. doi:10.1016/j.revpalbo.2020.104210. S2CID 216248477.
^ Yifan Hua; Xuelian Wang; Junlin Dong; Yanzhao Ji; Bainian Sun (2020). "A number of new seed fossils from the lower Permian of Gansu, Northwest China: implication for research on arils". Historical Biology: An International Journal of Paleobiology. 32 (8): 1098–1107. Bibcode:2020HBio...32.1098H. doi:10.1080/08912963.2019.1566323. S2CID 91993795.
^ ^a ^b Natalya Nosova (2020). "Female reproductive structures of Umaltolepis Krassilov and associated short shoots, buds and leaves of Pseudotorellia Florin from the Middle Jurassic of Angren, Uzbekistan". Review of Palaeobotany and Palynology. 281: Article 104266. Bibcode:2020RPaPa.28104266N. doi:10.1016/j.revpalbo.2020.104266. S2CID 219437430.
^ Josef Pšenička; Jun Wang; Jason Hilton; Weiming Zhou; Jiří Bek; Stanislav Opluštil; Jana Votočková Frojdová (2020). "A small heterophyllous vine climbing on Psaronius and Cordaites trees in the earliest Permian forests of North China" (PDF). International Journal of Plant Sciences. 181 (6): 616–645. doi:10.1086/708814. S2CID 219926230.
^ Weiming Zhou; Shan Wan; Mingli Wan; Jason Hilton; Josef Pšenička; Jun Wang (2020). "Yangopteris ascendens (Halle) gen. et comb. nov., a climbing alethopterid pteridosperm from the Asselian (earliest Permian) Wuda Tuff Flora". Review of Palaeobotany and Palynology. 294: Article 104282. doi:10.1016/j.revpalbo.2020.104282. S2CID 225462114.
^ Pedro Correia; Arden R. Bashforth; Zbynĕk Šimůnek; Christopher J. Cleal; Artur A. Sá; Conrad C. Labandeira (2020). "The history of herbivory on sphenophytes: a new calamitalean with an insect gall from the Upper Pennsylvanian of Portugal and a review of arthropod herbivory on an ancient lineage". International Journal of Plant Sciences. 181 (4): 387–418. doi:10.1086/707105. S2CID 214292984.
^ Ruiyun Li; Xiaoqiang Li; Xuelian Wang; Bainian Sun (2020). "First fossil liverwort with in situ flask-shaped receptacles from the Lower Cretaceous of Inner Mongolia, China". Cretaceous Research. 119: Article 104684. doi:10.1016/j.cretres.2020.104684. ISSN 0195-6671. S2CID 226314562.
^ Xiao-Yuan He; Shi-Jun Wang; Jason Hilton; Jean Galtier; Hong-Guan Jiang (2020). "An advanced species of the fern Botryopteris Renault from the Permian of southwestern China" (PDF). Review of Palaeobotany and Palynology. 273: Article 104136. Bibcode:2020RPaPa.27304136H. doi:10.1016/j.revpalbo.2019.104136. S2CID 210266014.
^ L. B. Golovneva; A. A. Grabovskiy; A. A. Zolina (2020). "A new species of the genus Birisia (Dicksoniaceae) from the lower–middle Albian deposits of Southern Primorye, Far East of Russia". Palaeobotany. 11: 74–95. doi:10.31111/palaeobotany/2020.11.74. S2CID 245112234.
^ Barry A. Thomas (2020). "A new species of leafy calamite stem from the Pennsylvanian (Bolsovian) of the South Wales Coalfield". Acta Palaeobotanica. 60 (1): 207–211. doi:10.35535/acpa-2020-0010. hdl:2160/cd78c623-2a50-4c65-a758-7207027deacd.
^ ^a ^b Yueyang Zhang; Hongxia Jiang; Yasheng Wu; Hongping Bao; Junfeng Ren; Zhengliang Huang (2020). "Calcified dasycladaleans from the Upper Ordovician in the Ordos Basin, China". Acta Micropalaeontologica Sinica. 37 (3): 228–237. doi:10.16087/j.cnki.1000-0674.2020.03.003.
^ ^a ^b ^c ^d Jordi Pérez-Cano; Telm Bover-Arnal; Carles Martín-Closas (2020). "Barremian charophytes from the Maestrat Basin". Cretaceous Research. 115. doi:10.1016/j.cretres.2020.104544. hdl:2445/171586. S2CID 224875084.
^ Jan Hinkelman; Lucia Vršanská (2020). "A Myanmar amber cockroach with protruding feces contains pollen and a rich microcenosis". The Science of Nature. 107 (2): Article number 13. Bibcode:2020SciNa.107...13H. doi:10.1007/s00114-020-1669-y. PMID 32125545. S2CID 211730431.
^ ^a ^b ^c Patricio Emmanuel Santamarina; Viviana Dora Barreda; Ari Iglesias; Augusto Nicolás Varela (2020). "Palynology from the Cenomanian Mata Amarilla Formation, southern Patagonia, Argentina". Cretaceous Research. 109: Article 104354. Bibcode:2020CrRes.10904354S. doi:10.1016/j.cretres.2019.104354. S2CID 212976442.
^ ^a ^b ^c ^d ^e ^f Ioan I. Bucur; Sylvain Rigaud; Nicolò Del Piero; Andrea Fucelli; Eric Heerwagen; Camille Peybernes; Giovan Peyrotty; Christian Verard; Jérôme Chablais; Rossana Martini (2020). "Upper Triassic calcareous algae from the Panthalassa Ocean". Rivista Italiana di Paleontologia e Stratigrafia. 126 (2): 499–540. doi:10.13130/2039-4942/13681.
^ Xiu-Cai Yuan; Cong-Hui Xiong; Fan-Kai Sun; Zi-Xi Wang; Teng Mao; Yi-Jie Li; Chun-Hui Liu; Ming-Xuan Sun; Jun-Ling Dong; Bai-Nian Sun (2020). "The geological significance of a new species of Coniopteris from the Middle Jurassic of northwestern China". Historical Biology: An International Journal of Paleobiology. 32 (2): 267–280. Bibcode:2020HBio...32..267Y. doi:10.1080/08912963.2018.1488251. S2CID 89752285.
^ Barry A. Thomas; Christopher J. Cleal (2020). "The nomenclature of fossil-taxa representing different preservational states: Lepidodendron as a case-study". Taxon. 69 (5): 1052–1061. doi:10.1002/tax.12291. S2CID 225235565.
^ Yang Yu; San-Ping Xie; John Devaney; Si-Hang Zhang; Tian-Yu Chen; Xu Zeng; Bing Wang; Yu Zhang (2020). "A new species of Drynaria (Polypodiaceae) from the late Miocene of Yunnan, Southwest China and implications on the genus evolution" (PDF). Palaeobiodiversity and Palaeoenvironments. 100 (4): 939–949. Bibcode:2020PdPe..100..939Y. doi:10.1007/s12549-020-00429-0. S2CID 219730737.
^ Li Zhang; Yong Wang; Hong-Yu Chen; Lei Han; Yu-Xin Zhang; Wen-Jia Li; Tao Yang; Hao-Jian Wang; Lin Bao; De-Fei Yan (2020). "New fossil material of Equicalastrobus (Equisetales) and associated leaves from the Late Triassic of Baojishan basin, Gansu Province, China". Historical Biology: An International Journal of Paleobiology. 33 (9): 1522–1533. doi:10.1080/08912963.2020.1716747. S2CID 213295920.
^ ^a ^b Aye Thida Aung; Jian Huang; Truong Van Do; Ai Song; Jia Liu; Zhe-Kun Zhou; Tao Su (2020). "Three new fossil records of Equisetum (Equisetaceae) from the Neogene of south-western China and northern Vietnam". PhytoKeys (138): 3–15. doi:10.3897/phytokeys.138.38674. PMC 6969027. PMID 31988601.
^ Branko Sokač; Tonći Grgasović (2020). "New dasycladalean alga with unusual two types of laterals from the Palaocene deposits of Konavle, SE of Dubrovnik (Dinarides, Croatia)". Revue de Micropaléontologie. 69: Article 100464. Bibcode:2020RvMic..6900464S. doi:10.1016/j.revmic.2020.100464. S2CID 226343559.
^ Ya Li; Yong-Dong Wang; Harald Schnerder; Peng-Cheng Wu (2020). "Frullania partita sp. nov. (Frullaniaceae, Porellales), a new leafy liverwort from the mid-Cretaceous of Myanmar". Cretaceous Research. 108: Article 104341. Bibcode:2020CrRes.10804341L. doi:10.1016/j.cretres.2019.104341. S2CID 213553976.
^ Yuriy S. Mamontov; John J. Atwood; Evgeny E. Perkovsky; Michael S. Ignatov (2020). "Hepatics from Rovno amber (Ukraine): Frullania pycnoclada and a new species, F. vanae". The Bryologist. 123 (3): 421–430. doi:10.1639/0007-2745-123.3.421. S2CID 225406886.
^ ^a ^b Fearghus R. McSweeney; Jeff Shimeta; John St. J. S. Buckeridge (2020). "Two new genera of early Tracheophyta (Zosterophyllaceae) from the upper Silurian–Lower Devonian of Victoria, Australia". Alcheringa: An Australasian Journal of Palaeontology. 44 (3): 379–396. Bibcode:2020Alch...44..379M. doi:10.1080/03115518.2020.1744725. S2CID 218993880.
^ Kelly C. Pfeiler; Alexandru M. F. Tomescu (2020). "An Early Devonian actinostelic euphyllophyte with secondary growth from the Emsian of Gaspé (Canada) and the importance of tracheid wall thickening patterns in early euphyllophyte systematics". Papers in Palaeontology. 7 (2): 1081–1095. doi:10.1002/spp2.1335. S2CID 225425418.
^ Weiming Zhou; Josef Pšenička; Jiří Bek; Mingli Wan; C. Kevin Boyce; Jun Wang (2020). "A new anachoropterid fern from the Asselian (Cisuralian) Wuda Tuff Flora". Review of Palaeobotany and Palynology. 294: Article 104346. doi:10.1016/j.revpalbo.2020.104346. S2CID 228855095.
^ Steven T. LoDuca; Anthony L. Swinehart; Matthew A. LeRoy; Denis K. Tetreault; Shawn Steckenfinger (2020). "Codium-like taxa from the Silurian of North America: morphology, taxonomy, paleoecology, and phylogenetic affinity". Journal of Paleontology. 95 (2): 207–235. doi:10.1017/jpa.2020.85. S2CID 228938523.
^ Li-jun Chen; Ye-mao Hou; Peng-fei Yin; Xin Wang (2020). "An edible fruit from the Jurassic of China". China Geology. 3 (1): 8–15. Bibcode:2020CGeo....3....8C. doi:10.31035/cg2020010.
^ Antoine Champreux; Brigitte Meyer-Berthaud; Anne-Laure Decombeix (2020). "Keraphyton gen. nov., a new Late Devonian fern-like plant from Australia". PeerJ. 8: e9321. doi:10.7717/peerj.9321. PMC 7304422. PMID 32587800.
^ D. S. Kopylov; A. P. Rasnitsyn; D. S. Aristov; A. S. Bashkuev; N. V. Bazhenova; V. Yu. Dmitriev; A. V. Gorochov; M. S. Ignatov; V. D. Ivanov; A. V. Khramov; A. A. Legalov; E. D. Lukashevich; Yu. S. Mamontov; S. I. Melnitsky; B. Ogłaza; A. G. Ponomarenko; A. A. Prokin; O. V. Ryzhkova; A. S. Shmakov; N. D. Sinitshenkova; A. Yu. Solodovnikov; O. D. Strelnikova; I. D. Sukacheva; A. V. Uliakhin; D. V. Vasilenko; P. Wegierek; E. V. Yan; M. Zmarzły (2020). "The Khasurty Fossil Insect Lagerstätte". Paleontological Journal. 54 (11): 1221–1394. Bibcode:2020PalJ...54.1221K. doi:10.1134/S0031030120110027. S2CID 231850225.
^ Yu.V. Mosseichik (2020). "Lepidodendron species from the Viséan of the Moscow Basin" (PDF). Lethaea Rossica. 20: 19–33.
^ Yi-zhi Xu; Ning Yue; Shi Liu; Yi Zhang; Shao-lin Zheng; Yong-dong Wang; Xin-ran Huo (2020). "Lobatannularia linjiaensis sp. nov. from the Middle Triassic of Benxi, Liaoning". Acta Palaeontologica Sinica. 59 (3): 329–344. doi:10.19800/j.cnki.aps.2019.024.
^ Sergey V. Naugolnykh; Ming Tu; Xiao-Yan Liu; Jian-Hua Jin (2020). "A new species of Lygodium (Schizaeaceae) from the Buxin Formation (Paleocene), Sanshui Basin, South China". Palaeoworld. 29 (3): 606–616. doi:10.1016/j.palwor.2019.07.003. S2CID 199101009.
^ Tomoyuki Katagiri; Hisao Shinden (2020). "Discovery of a simple thalloid liverwort Metzgeriites kujiensis (Metzgeriaceae), a new species from Late Cretaceous Japanese amber". Hattoria. 11: 13–21. doi:10.18968/hattoria.11.0_13.
^ Silvia Gnaedinger; Bárbara Cariglino; Ana María Zavattieri; Mariana Monti; Pedro R. Gutiérrez (2020). "Neoarthropitys gondwanaensis gen. et sp. nov. from the Middle Triassic of Gondwana: An intermediate stage in the anatomical trend of Equisetalean stems". Review of Palaeobotany and Palynology. 282: Article 104298. Bibcode:2020RPaPa.28204298G. doi:10.1016/j.revpalbo.2020.104298. S2CID 225347405.
^ ^a ^b Yeming Cheng; Fengxiang Liu; Xiaonan Yang; Tongxing Sun (2020). "Two new species of Mesozoic tree ferns (Osmundaceae: Osmundacaulis) in Eurasia as evidence of long-term geographic isolation". Geoscience Frontiers. 11 (5): 1875–1888. Bibcode:2020GeoFr..11.1875C. doi:10.1016/j.gsf.2020.01.019.
^ ^a ^b Ana M. Zavattieri; Pedro R. Gutiérrez; Mariana Monti (2020). "Middle Triassic freshwater green algae and fungi of the Puesto Viejo Basin, central-western Argentina: palaeoenvironmental implications". Alcheringa: An Australasian Journal of Palaeontology. 44 (3): 430–459. Bibcode:2020Alch...44..430Z. doi:10.1080/03115518.2020.1749302. S2CID 219519003.
^ Li Liu; Josef Pšenička; Jiří Bek; Mingli Wan; Hermann W. Pfefferkorn; Jun Wang (2020). "A whole calamitacean plant Palaeostachya guanglongii from the Asselian (Permian) Taiyuan Formation in the Wuda Coalfield, Inner Mongolia, China". Review of Palaeobotany and Palynology. 294: Article 104245. doi:10.1016/j.revpalbo.2020.104245. S2CID 219017989.
^ Ruiyun Li; Xiaoqiang Li; Shenghui Deng; Bainian Sun (2020). "Morphology and microstructure of Pellites hamiensis nov. sp., a Middle Jurassic liverwort from northwestern China and its evolutionary significance". Geobios. 62: 23–29. Bibcode:2020Geobi..62...23L. doi:10.1016/j.geobios.2020.07.003. S2CID 225500594.
^ P. Hiller; M. Böhme; S. Schneider; J. Prieto; B. Bomfleur (2020). "Plenasium (Aurealcaulis) elegans sp. nov. from the Eocene of Vietnam – a connecting link in the evolution of modern Royal Ferns (Osmundeae, Osmundaceae)". Journal of Systematic Palaeontology. 18 (8): 703–715. Bibcode:2020JSPal..18..703H. doi:10.1080/14772019.2019.1683771. S2CID 209573114.
^ Yi Zhang; Yong-Dong Wang; Yue Hong; Liu Cao; Fu-liang Gao (2020). "Pleuromeia discovered from the Middle Triassic Linjia Formation of Benxi, Northeast China". Palaeoworld. 29 (4): 706–714. doi:10.1016/j.palwor.2020.02.004. S2CID 216475501.
^ Carmen Álvarez-Vázquez; Jiří Bek; Jana Drábková (2020). "Polysporia baetica sp. nov., a new heterosporous sub-arborescent isoetalean from lower Bolsovian (Middle Pennsylvanian) strata of the Peñarroya-Belmez-Espiel Coalfield (Córdoba, SW Spain)". Review of Palaeobotany and Palynology. 272: Article 104115. Bibcode:2020RPaPa.27204115A. doi:10.1016/j.revpalbo.2019.104115.
^ Zlatko Kvaček; Vasilis Teodoridis (2020). "A new Oligocene fern of Dryopteridaceae from the Českéstředohoří Mts (Czech Republic)". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 295 (1): 9–16. doi:10.1127/njgpa/2020/0864. S2CID 213778635.
^ Chunxiang Li; Robbin C. Moran; Junye Ma; Bo Wang; Jiasheng Hao (2020). "A new fossil record of Lindsaeaceae (Polypodiales) from the mid-Cretaceous amber of Myanmar". Cretaceous Research. 105: Article 104040. Bibcode:2020CrRes.10504040L. doi:10.1016/j.cretres.2018.12.010.
^ Qing Tang; Ke Pang; Xunlai Yuan; Shuhai Xiao (2020). "A one-billion-year-old multicellular chlorophyte". Nature Ecology & Evolution. 4 (4): 543–549. Bibcode:2020NatEE...4..543T. doi:10.1038/s41559-020-1122-9. PMC 8668152. PMID 32094536. S2CID 211266015.
^ ^a ^b Josep Sanjuan; Ingeborg Soulié-Märsche (2020). "New charophyte assemblage from middle Miocene lacustrine deposits of Moneva (Ebro Basin, Spain)". Geobios. 59: 79–90. Bibcode:2020Geobi..59...79S. doi:10.1016/j.geobios.2020.03.001. S2CID 218775725.
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[1] Shook Ling Low; Tao Su; Teresa E. V. Spicer; Fei-Xiang Wu; Tao Deng; Yao-Wu Xing; Zhe-Kun Zhou (2020). "Oligocene Limnobiophyllum (Araceae) from the central Tibetan Plateau and its evolutionary and palaeoenvironmental implications". Journal of Systematic Palaeontology. 18 (5): 415–431. Bibcode:2020JSPal..18..415L. doi:10.1080/14772019.2019.1611673. S2CID 208589882.

[PE233Naetal-2] Yuling Na; Jane Blanchard; Hongshan Wang (2020). "Fruits, seeds and flowers from the Puryear clay pit (middle Eocene Cockfield Formation), western Tennessee, USA". Palaeontologia Electronica. 23 (3): Article number 23(3):a49. doi:10.26879/1045.

[Scabrastephanoporites-3] Vann Smith; Sophie Warny; David M. Jarzen; Thomas Demchuk; Vivi Vajda; Sean P.S. Gulick (2020). "Paleocene–Eocene palynomorphs from the Chicxulub impact crater, Mexico. Part 2: angiosperm pollen". Palynology. 44 (3): 489–519. Bibcode:2020Paly...44..489S. doi:10.1080/01916122.2019.1705417. S2CID 213827225.

[4] Mahasin Ali Khan; Manoshi Hazra; Sumana Mahato; Robert A. Spicer; Kaustav Roy; Taposhi Hazra; Manosij Bandopadhaya; Teresa E.V. Spicer; Subir Bera (2020). "A Cretaceous Gondwana origin of the wax palm subfamily (Ceroxyloideae: Arecaceae) and its paleobiogeographic context". Review of Palaeobotany and Palynology. 283: Article 104318. Bibcode:2020RPaPa.28304318K. doi:10.1016/j.revpalbo.2020.104318. S2CID 224946279.

[5] Mahasin Ali Khan; Kaustav Roy; Taposhi Hazra; Sumana Mahato; Subir Bera (2020). "A new coryphoid palm from the Maastrichtian-Danian sediments of Madhya Pradesh and its palaeoenvironmental implications". Journal of the Geological Society of India. 95 (1): 75–83. Bibcode:2020JGSI...95...75K. doi:10.1007/s12594-020-1388-1. S2CID 210134584.

[6] David Robert Greenwood; John G. Conran (2020). "Fossil coryphoid palms from the Eocene of Vancouver, British Columbia, Canada". International Journal of Plant Sciences. 181 (2): 224–240. doi:10.1086/706450. S2CID 208587364.

[7] Kaustav Roy; Taposhi Hazra; Manoshi Hazra; Sumana Mahato; Subir Bera; Mahasin Ali Khan (2020). "A new coryphoid costapalmate palm leaf from the Maastrichtian-Danian of India". Botany Letters. 168 (2): 155–166. doi:10.1080/23818107.2020.1845974. S2CID 229408918.

[8] Patricia Vallati; Andrea De Sosa Tomas; Gabriel Casal (2020). "A Maastrichtian terrestrial palaeoenvironment close to the K/Pg boundary in the Golfo San Jorge basin, Patagonia, Argentina". Journal of South American Earth Sciences. 97: Article 102401. Bibcode:2020JSAES..9702401V. doi:10.1016/j.jsames.2019.102401.

[PPKvaceketal-9] ^ ^a ^b ^c ^d ^e ^f ^g ^h Zlatko Kvaček; Vasilis Teodoridis; Thomas Denk (2020). "The Pliocene flora of Frankfurt am Main, Germany: taxonomy, palaeoenvironments and biogeographic affinities". Palaeobiodiversity and Palaeoenvironments. 100 (3): 647–703. Bibcode:2020PdPe..100..647K. doi:10.1007/s12549-019-00391-6.

[Palynology451-10] ^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k ^l ^m ⁿ ^o ^p ^q ^r ^s ^t ^u ^v ^w ^x ^y ^z ^aa ^ab ^ac ^ad ^ae ^af ^ag ^ah ^ai ^aj Fátima Praxedes Rabelo Leite; Silane Aparecida Ferreira da Silva-Caminha; Carlos D'Apolito (2020). "New Neogene index pollen and spore taxa from the Solimões Basin (Western Amazonia), Brazil". Palynology. 45 (1): 115–141. doi:10.1080/01916122.2020.1758971. S2CID 219090032.

[11] M. Laura Pipo; Ari Iglesias; Josefina Bodnar (2020). "A new vesselless angiosperm stem with a cambial variant from the Upper Cretaceous of Antarctica". Acta Palaeontologica Polonica. 65 (2): 261–272. doi:10.4202/app.00697.2019. hdl:11336/136234.

[IJPSHuegeleManchester-12] Indah B. Huegele; Steven R. Manchester (2020). "An early Paleocene carpoflora from the Denver Basin of Colorado, USA, and its implications for plant-animal interactions and fruit size evolution". International Journal of Plant Sciences. 181 (6): 646–665. doi:10.1086/707727. S2CID 219748982.

[AP601Hably-13] Lilla Hably (2020). "The Karpatian (late early Miocene) flora of the Mecsek area". Acta Palaeobotanica. 60 (1): 51–122. doi:10.35535/acpa-2020-0003.

[14] Thomas Denk; Johannes Martin Bouchal; Pavel Smirnov; Yaroslav Trubin (2020). "Late Oligocene leaf and pollen flora of Southwestern Siberia: taxonomy, biogeography and palaeoenvironments". Historical Biology: An International Journal of Paleobiology. 33 (11): 2951–2976. doi:10.1080/08912963.2020.1839064. S2CID 230547821.

[15] Hai Zhu; Steven R. Manchester (2020). "Fruit of Staphylea (Staphyleaceae) from the Oligocene of Montana, USA". Review of Palaeobotany and Palynology. 280: Article 104275. Bibcode:2020RPaPa.28004275Z. doi:10.1016/j.revpalbo.2020.104275. S2CID 225729365.

[16] Susanne S. Renner; Viviana D. Barreda; María Cristina Tellería; Luis Palazzesi; Tanja M. Schuster (2020). "Early evolution of Coriariaceae (Cucurbitales) in light of a new early Campanian (ca. 82 Mya) pollen record from Antarctica". Taxon. 69 (1): 87–99. doi:10.1002/tax.12203.

[17] MacKenzie Allan Smith; Steven R. Manchester (2020). "CT-scans of capsules from the Clarno Formation (Oregon, USA) reveal an extinct Eocene theaceous taxon". Acta Palaeobotanica. 60 (2): 251–258. doi:10.35535/acpa-2020-0013.

[18] Xiang-Chuan Li; Steven R. Manchester; Qin Wang; Liang Xiao; Tian-Long Qi; Yun-Zhi Yao; Dong Ren; Qiang Yang (2020). "A unique record of Cercis from the late early Miocene of interior Asia and its significance for paleoenvironment and paleophytogeography". Journal of Systematics and Evolution. 59 (6): 1321–1338. doi:10.1111/jse.12640. S2CID 219523602.

[Bastardioxylon-19] Johanna Baez; Alexandra Crisafulli (2020). "Novelties in the xylotaphoflora from Chiquimil Formation (Miocene), Catamarca-Argentina". Journal of South American Earth Sciences. 107: Article 102943. doi:10.1016/j.jsames.2020.102943. S2CID 225109309.

[Veraguasoxylon-20] Oris J. Rodríguez-Reyes; Emilio Estrada-Ruiz (2020). "Two new reports of ancient rainforest trees from the Azuero Peninsula, Panama". Ameghiniana. 57 (3): 209–218. doi:10.5710/AMGH.22.02.2020.3299. S2CID 216250364.

[BZ1051-21] A.L. Averyanova; Yaowu Xing (2020). "New Paleogene angiosperm species of Zaissan Depression (eastern Kazakhstan)". Botanicheskii Zhurnal. 105 (1): 46–57. doi:10.31857/S0006813620010044. S2CID 218793001.

[FEEFagaceae-22] ^ ^a ^b ^c ^d ^e ^f ^g ^h Xiaoyan Liu; Hanzhang Song; Jianhua Jin (2020). "Diversity of Fagaceae on Hainan Island of South China during the middle Eocene: implications for phytogeography and paleoecology". Frontiers in Ecology and Evolution. 8: Article 255. doi:10.3389/fevo.2020.00255. S2CID 221117336.

[PDCarpinus-23] Li Xue; Linbo Jia; Gi-soo Nam; Yongjiang Huang; Shitao Zhang; Yuqing Wang; Zhuo Zhou; Yongsheng Chen (2020). "Involucre fossils of Carpinus, a northern temperate element, from the Miocene of China and the evolution of its species diversity in East Asia". Plant Diversity. 42 (3): 155–167. doi:10.1016/j.pld.2020.01.001. PMC 7361179. PMID 32695948.

[24] Anthony L. Swinehart; James O. Farlow (2020). "Plant and invertebrate macrofossils from the Pipe Creek Sinkhole (Late Neogene), Grant County, Indiana". Historical Biology: An International Journal of Paleobiology. 33 (11): 3111–3140. doi:10.1080/08912963.2020.1851686. S2CID 230538832.

[25] Dimitra Mantzouka; Miroslav Ivanov; Vladimir Bozukov (2020). "The first report of an 'evergreen Castanopsis type' wood (Fagaceae) for the Late Miocene–Early Pliocene of Europe (Bulgaria, Blagoevgrad Graben)". Geological Journal. 56 (2): 786–803. doi:10.1002/gj.3919.

[Willdenowia-26] ^ ^a ^b ^c ^d ^e ^f Eva-Maria Sadowski; Alexander R. Schmidt; Thomas Denk (2020). "Staminate inflorescences with in situ pollen from Eocene Baltic amber reveal high diversity in Fagaceae (oak family)". Willdenowia. 50 (3): 405–517. doi:10.3372/wi.50.50303.

[27] Cédric Del Rio; Teng-Xiang Wang; Jia Liu; Shui-Qing Liang; Robert A. Spicer; Fei-Xiang Wu; Zhe-Kun Zhou; Tao Su (2020). "Asclepiadospermum gen. nov., the earliest fossil record of Asclepiadoideae (Apocynaceae) from the early Eocene of central Qinghai-Tibetan Plateau, and its biogeographic implications". American Journal of Botany. 107 (1): 126–138. doi:10.1002/ajb2.1418. PMID 31944266.

[28] Cédric Del Rio; Jian Huang; Gregory W. Stull; Rémi Allemand; Zhe-Kun Zhou; Tao Su (2020). "First macrofossil record of Icacinaceae in East Asia (early Oligocene, Wenshan Basin) and its ecological implications". Journal of Systematics and Evolution. 60 (2): 445–455. doi:10.1111/jse.12700. S2CID 228976920.

[29] Cédric Del Rio; Dario De Franceschi (2020). "Icacinaceae fossil fruits from three sites of the Paris Basin (early Eocene, France): local diversity and global biogeographic implications". Geodiversitas. 42 (2): 17–28. doi:10.5252/geodiversitas2020v42a2.^{[permanent dead link]}

[30] Andrew C. Rozefelds; Gregory Stull; Peta Hayes; David R. Greenwood (2020). "The fossil record of Icacinaceae in Australia supports long-standing Palaeo-Antarctic rainforest connections in southern high latitudes". Historical Biology: An International Journal of Paleobiology. 33 (11): 2854–2864. doi:10.1080/08912963.2020.1832089. S2CID 229005088.

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[32] Ünal Akkemik; Dimitra Mantzouka; Umut Tunç; Fikret Koçbulut (2020). "The first paleoxylotomical evidence from the Mid-Eocene Climate Optimum from Turkey". Review of Palaeobotany and Palynology. 285: Article 104356. doi:10.1016/j.revpalbo.2020.104356. S2CID 229395041.

[Patagonoxylon-33] Daniela P. Ruiz; M. Sol Raigemborn; Mariana Brea; Roberto R. Pujana (2020). "Paleocene Las Violetas Fossil Forest: Wood anatomy and paleoclimatology". Journal of South American Earth Sciences. 98: Article 102414. Bibcode:2020JSAES..9802414R. doi:10.1016/j.jsames.2019.102414. S2CID 213796947.

[34] Kenton L. Chambers; George O. Poinar, Jr (2020). "Thymolepis toxandra gen. et sp. nov., a mid-Cretaceous fossil flower with horseshoe-shaped anthers". Journal of the Botanical Research Institute of Texas. 14 (1): 57–64. doi:10.17348/jbrit.v14.i1.896.

[35] George O. Poinar, Jr; Kenton L. Chambers; Urszula T. Iwaniec; Fernando E. Vega (2020). "Valviloculus pleristaminis gen. et sp. nov., a Lauralean fossil flower with valvate anthers from mid-Cretaceous Myanmar amber". Journal of the Botanical Research Institute of Texas. 14 (2): 359–366. doi:10.17348/jbrit.v14.i2.1014.

[GJSmilax-36] Jun-Ling Dong; Bai-Nian Sun; Ai-Jing Li; Hui Chen (2020). "The diversity of Smilax (Smilacaceae) leaves from the Middle Miocene in southeastern China". Geological Journal. 56 (2): 744–757. doi:10.1002/gj.3882. S2CID 225790148.

[37] Lu-Liang Huang; Jian-Hua Jin; Cheng Quan; Alexei A. Oskolski (2020). "Mummified Magnoliaceae woods from the upper Oligocene of South China, with biogeography, paleoecology, and wood trait evolution implications". Journal of Systematics and Evolution. 58 (1): 89–100. doi:10.1111/jse.12480. S2CID 91861708.

[38] Else Marie Friis; Peter R. Crane; Kaj Raunsgaard Pedersen (2020). "Melloniflora, a new extinct multiparted flower from the Early Cretaceous of Virginia, USA". International Journal of Plant Sciences. 181 (9): 887–897. doi:10.1086/710490. S2CID 224837365.

[39] Atsufumi Narita; Atsushi Yabe; Kazuhiko Uemura; Midori Matsumoto (2020). "Late middle Miocene Konan flora from northern Hokkaido, Japan". Acta Palaeobotanica. 60 (2): 259–295. doi:10.35535/acpa-2020-0012.

[40] Mahasin Ali Khan; Robert A. Spicer; Teresa E. V. Spicer; Kaustav Roy; Manoshi Hazra; Taposhi Hazra; Sumana Mahato; Sanchita Kumar; Subir Bera (2020). "Dipterocarpus (Dipterocarpaceae) leaves from the K-Pg of India: a Cretaceous Gondwana presence of the Dipterocarpaceae". Plant Systematics and Evolution. 306 (6): Article 90. Bibcode:2020PSyEv.306...90K. doi:10.1007/s00606-020-01718-z. S2CID 228870254.

[41] Maria C. Zamaloa; Maria A. Gandolfo; Kevin C. Nixon (2020). "52 million years old Eucalyptus flower sheds more than pollen grains". American Journal of Botany. 107 (12): 1763–1771. doi:10.1002/ajb2.1569. PMC 7839439. PMID 33274448.

[42] Eliana Moya; Mariana Brea (2020). "Combretaceous fossil wood from Ituzaingó Formation (Late Miocene?), Argentina, indicate a coastal marine environment". Review of Palaeobotany and Palynology. 281: Article 104270. Bibcode:2020RPaPa.28104270M. doi:10.1016/j.revpalbo.2020.104270. hdl:11336/142562. S2CID 224947789.

[43] Ya Li; Yi-Ming Cui; Carole T. Gee; Xiao-Qing Liang; Cheng-Sen Li (2020). "Primotrapa gen. nov., an extinct transitional genus bridging the evolutionary gap between Lythraceae and Trapoideae, from the early Miocene of North China". BMC Evolutionary Biology. 20 (1): 150. Bibcode:2020BMCEE..20..150L. doi:10.1186/s12862-020-01697-2. PMC 7661254. PMID 33183234.

[44] David Peris; Conrad C. Labandeira; Eduardo Barrón; Xavier Delclòs; Jes Rust; Bo Wang (2020). "Generalist pollen-feeding beetles during the mid-Cretaceous". iScience. 23 (3): Article 100913. Bibcode:2020iSci...23j0913P. doi:10.1016/j.isci.2020.100913. PMC 7113562. PMID 32191877.

[45] Nathan A. Jud; Maria A. Gandolfo (2020). "Fossil evidence from South America for the diversification of Cunoniaceae by the earliest Palaeocene". Annals of Botany. 127 (3): 305–315. doi:10.1093/aob/mcaa154. PMC 7872129. PMID 32860407.

[JSEElaeocarpus-46] ^ ^a ^b ^c ^d ^e ^f Xiao-Yan Liu; Steven R. Manchester; Andrew C. Rozefelds; Cheng Quan; Jian-Hua Jin (2020). "First fossil fruits of Elaeocarpus (Elaeocarpaceae) in East Asia: implications for phytogeography and paleoecology". Journal of Systematics and Evolution. 60 (2): 456–471. doi:10.1111/jse.12684. S2CID 234429843.

[Circinites-47] Juan M. Robledo; Luisa M. Anzótegui; Olga G. Martínez; Ricardo N. Alonso (2020). "Flora and insect trace fossils from the Mio-Pliocene Quebrada del Toro locality (Gobernador Solá, Salta, Argentina)". Journal of South American Earth Sciences. 100: Article 102544. Bibcode:2020JSAES.10002544R. doi:10.1016/j.jsames.2020.102544. S2CID 216377729.

[AJBBanksia-48] Raymond J. Carpenter; Lynne A. Milne (2020). "New species of xeromorphic Banksia (Proteaceae) foliage and Banksia-like pollen from the late Eocene of Western Australia". Australian Journal of Botany. 68 (3): 165–178. doi:10.1071/BT19110. S2CID 214113278.

[49] Indah Badriyyah Huegele; Robert J. Spielbauer; Steven R. Manchester (2020). "Morphology and systematic affinities of Platanus dissecta Lesquereux (Platanaceae) from the Miocene of western North America". International Journal of Plant Sciences. 181 (3): 324–341. doi:10.1086/706453. S2CID 208566485.

[DelRioetalMenispermaceae-50] ^ ^a ^b ^c ^d ^e ^f Cédric Del Rio; Jian Huang; Ping Liu; Wei-Yu-Dong Deng; Teresa E.V. Spicer; Fei-Xiang Wu; Zhe-Kun Zhou; Tao Su (2020). "New Eocene fossil fruits and leaves of Menispermaceae from the central Tibetan Plateau and their biogeographic implications". Journal of Systematics and Evolution. 59 (6): 1287–1306. doi:10.1111/jse.12701. S2CID 228951072.

[51] Hui Jia; David K. Ferguson; Bainian Sun; Xiangning Meng; Yifan Hua (2020). "Phytogeographic implications of a fossil endocarp of Diploclisia (Menispermaceae) from the Miocene of eastern China". Geological Journal. 56 (2): 758–767. doi:10.1002/gj.3867. S2CID 219907004.

[Paleoorbicarpum-52] Meng Han; Xin-Kai Wu; Ming Tu; Tatiana M. Kodrul; Jian-Hua Jin (2020). "Diversity of Menispermaceae from the Paleocene and Eocene of South China". Journal of Systematics and Evolution. 58 (3): 354–366. doi:10.1111/jse.12499. S2CID 199062171.

[53] Zhekun Zhou; Tengxiang Wang; Jian Huang; Jia Liu; Weiyudong Deng; Shihu Li; Chenglong Deng; Tao Su (2020). "Fossil leaves of Berhamniphyllum (Rhamnaceae) from Markam, Tibet and their biogeographic implications". Science China Earth Sciences. 63 (2): 224–234. Bibcode:2020ScChD..63..224Z. doi:10.1007/s11430-019-9477-8. S2CID 211028504.

[54] Zixi Wang; Fabiany Herrera; Junwu Shu; Suxin Yin; Gongle Shi (2020). "A new Choerospondias (Anacardiaceae) endocarp from the middle Miocene of Southeast China and its paleoecological implications". Review of Palaeobotany and Palynology. 283: Article 104312. Bibcode:2020RPaPa.28304312W. doi:10.1016/j.revpalbo.2020.104312.

[55] Oris Rodríguez-Reyes; Emilio Estrada-Ruiz; Peter Gasson (2020). "Evidence of large Anacardiaceae trees from the Oligocene–early Miocene Santiago Formation, Azuero, Panama". Boletín de la Sociedad Geológica Mexicana. 72 (2): Article A300719. doi:10.18268/BSGM2020v72n2a300719.

[56] Brian A. Atkinson (2020). "Fossil evidence for a Cretaceous rise of the mahogany family". American Journal of Botany. 107 (1): 139–147. doi:10.1002/ajb2.1416. PMID 31903551.

[57] M. Jimena Franco; Eliana Moya; Mariana Brea; Camila Martínez Martínez (2020). "Astroniumxylon, Schinopsixylon, and Parametopioxylon n. gen. fossil woods from upper Cenozoic of Argentina: taxonomic revision, new taxon and new records". Journal of Paleontology. 94 (2): 185–201. Bibcode:2020JPal...94..185F. doi:10.1017/jpa.2019.97. S2CID 212737596.

[58] Steven R. Manchester; Kory A. Disney; Kasey K. Pham (2020). "Winged fruits of rutaceous affinity from the Eocene of western North America". Fossil Imprint. 76 (2): 211–216. doi:10.37520/fi.2020.018.

[59] Zack J. Quirk; Elizabeth J. Hermsen (2020). "Neogene Corylopsis Seeds from Eastern Tennessee". Journal of Systematics and Evolution. 59 (3): 611–621. doi:10.1111/jse.12571. S2CID 214264053.

[60] Yi-Min Tian; Jian Huang; Tao Su; Shi-Tao Zhang (2020). "Early Oligocene Itea (Iteaceae) leaves from East Asia and their biogeographic implications". Plant Diversity. 43 (2): 142–151. doi:10.1016/j.pld.2020.09.006. PMC 8103422. PMID 33997547.

[61] Alex R. Scharfstein; Ruth A. Stockey; Gar W. Rothwell (2020). "Evolution and phylogeny of Altingiaceae: anatomically preserved infructescences from Late Cretaceous deposits of Vancouver Island, British Columbia, Canada". International Journal of Plant Sciences. 181 (4): 452–463. doi:10.1086/707107. S2CID 216212606.

[62] Rocío Deanna; Peter Wilf; Maria A. Gandolfo (2020). "New physaloid fruit-fossil species from early Eocene South America". American Journal of Botany. 107 (12): 1749–1762. doi:10.1002/ajb2.1565. PMID 33247843. S2CID 227192001.

[Eotrochion-63] Steven R. Manchester; Zlatko Kvaček; Walter S. Judd (2020). "Morphology, anatomy, phylogenetics and distribution of fossil and extant Trochodendraceae in the Northern Hemisphere". Botanical Journal of the Linnean Society. 195 (3): 467–484. doi:10.1093/botlinnean/boaa046.

[64] Wen-Long He; Xiao-Jing Wang (2020). "A Miocene flora from the Toupi Formation in Jiangxi Province, southeastern China". Palaeoworld. 30 (4): 757–769. doi:10.1016/j.palwor.2020.12.006. S2CID 234387044.

[65] Ezequiel Ignacio Vera; Valeria S. Perez Loinaze; Magdalena Llorens; Mauro Gabriel Passalia (2020). "The fossil genus Aextoxicoxylon (Magnoliopsida) in the Upper Cretaceous Puntudo Chico Formation, Chubut Province, Argentina". Cretaceous Research. 107: Article 104315. Bibcode:2020CrRes.10704315V. doi:10.1016/j.cretres.2019.104315. S2CID 210254812.

[Atlantocarpus-66] Else Marie Friis; Peter R. Crane; Kaj Raunsgaard Pedersen (2020). "Multiparted, apocarpous flowers from the Early Cretaceous of eastern North America and Portugal". Fossil Imprint. 76 (2): 279–296. doi:10.37520/fi.2020.023.

[67] Else Marie Friis; Peter R. Crane; Kaj Raunsgaard Pedersen (2020). "Catanthus, An Extinct Magnoliid Flower From The Early Cretaceous Of Portugal". International Journal of Plant Sciences. 182 (1): 28–45. doi:10.1086/711081. S2CID 228939581.

[68] Caroline Siegert; Elizabeth J. Hermsen (2020). "Cavilignum pratchettii gen. et sp. nov., a novel type of fossil endocarp with open locules from the Neogene Gray Fossil Site, Tennessee, USA". Review of Palaeobotany and Palynology. 275: Article 104174. Bibcode:2020RPaPa.27504174S. doi:10.1016/j.revpalbo.2020.104174. S2CID 213393197.

[69] George O. Poinar, Jr; Kenton L. Chambers (2020). "Chainandra zeugostylus gen. et sp. nov., a mid-Cretaceous amber fossil with sagittate anthers opening widely at maturity". Journal of the Botanical Research Institute of Texas. 14 (2): 367–372. doi:10.17348/jbrit.v14.i2.1015.

[70] George O. Poinar, Jr; Kenton L. Chambers (2020). "Cyathitepala papillosa gen. et sp. nov., a mid-Cretaceous fossil flower from Myanmar amber with valvate anthers". Journal of the Botanical Research Institute of Texas. 14 (2): 351–358. doi:10.17348/jbrit.v14.i2.1013.

[71] George O. Poinar, Jr; Kenton L. Chambers (2020). "Dasykothon leptomiscus gen. et sp. nov., a fossil flower of possible Lauralean affinity from Myanmar amber". Journal of the Botanical Research Institute of Texas. 14 (1): 65–71. doi:10.17348/jbrit.v14.i1.897.

[72] Xue-Die Liu; José Bienvenido Diez; Yong Fan; Zhong-Jian Liu; Xin Wang (2020). "A unique flower in Miocene amber sheds new light on the evolution of flowers". Palaeoentomology. 3 (4): 423–432. doi:10.11646/palaeoentomology.3.4.15.

[73] Gang Han; Xin Wang (2020). "A New Infructescence of Angiosperms from the Early Cretaceous of China". Acta Geologica Sinica (English Edition). 94 (5): 1711–1713. Bibcode:2020AcGlS..94.1711H. doi:10.1111/1755-6724.14591. S2CID 225166235.

[74] Dieter Uhl; Khum N. Paudayal; Sophie Hervet; Haytham El Atfy (2020). "Menatanthus mosbruggeri gen. nov. et sp. nov. – A flower with in situ pollen tetrads from the Paleocene maar lake of Menat (Puy-de-Dôme, France)". Palaeobiodiversity and Palaeoenvironments. 101 (1): 51–58. doi:10.1007/s12549-020-00453-0. S2CID 226960216.

[75] George O. Poinar, Jr; Kenton L. Chambers (2020). "Phantophlebia dicycla gen. et sp. nov., a five-merous fossil flower in mid-Cretaceous Myanmar amber". Journal of the Botanical Research Institute of Texas. 14 (1): 73–80. doi:10.17348/jbrit.v14.i1.898.

[76] Adam T. Halamski; Jiří Kvaček; Marcela Svobodová; Ewa Durska; Zuzana Heřmanová (2020). "Late Cretaceous mega-, meso-, and microfloras from Lower Silesia". Acta Palaeontologica Polonica. 65 (4): 811–878. doi:10.4202/app.00744.2020.

[77] Lida Xing; Lei Gu (2020). "The possible earliest epizoochorous fruit preserved in mid-Cretaceous Burmese amber". Cretaceous Research. 114: Article 104498. Bibcode:2020CrRes.11404498X. doi:10.1016/j.cretres.2020.104498. S2CID 219439641.

[78] Deepak D. Ramteke; Steven R. Manchester; Vaishali D. Nagrale; Selena Y. Smith (2020). "Singpuria, a new genus of Eudicot flower from the latest Cretaceous Deccan Intertrappean Beds of India". Acta Palaeobotanica. 60 (2): 323–332. doi:10.35535/acpa-2020-0017.

[79] Zhong-Jian Liu; Li-Jun Chen; Xin Wang (2020). "A whole-plant monocot from the Lower Cretaceous". Palaeoworld. 30 (1): 169–175. doi:10.1016/j.palwor.2020.03.008.

[80] Xuedie Liu; Liang Ma; Bin Liu; Zhong-Jian Liu; Xin Wang (2020). "A novel angiosperm including various parts from the Early Cretaceous sheds new light on flower evolution". Historical Biology: An International Journal of Paleobiology. 33 (11): 2706–2714. doi:10.1080/08912963.2020.1825411.

[81] Xiaoqing Zhang; Yongdong Wang; David L. Dilcher; Steven R. Manchester (2020). "Wireroadia, a new genus of winged fruit from the Cretaceous of Alabama and New England, USA". International Journal of Plant Sciences. 181 (9): 898–910. doi:10.1086/710492. S2CID 224836733.

[82] Alexei A. Oskolski; Luliang Huang; Anna V. Stepanova; Jianhua Jin (2020). "Araucarioid wood from the late Oligocene–early Miocene of Hainan Island: first fossil evidence for the genus Agathis in the Northern Hemisphere". Journal of Plant Research. 133 (2): 157–173. Bibcode:2020JPlR..133..157O. doi:10.1007/s10265-019-01165-z. PMID 31915952. S2CID 210088380.

[JPTronquimalalGroup-83] Silvia C. Gnaedinger; Ana María Zavattieri (2020). "Coniferous woods from the Upper Triassic of southwestern Gondwana, Tronquimalal Group, Neuquén Basin, Mendoza Province, Argentina". Journal of Paleontology. 94 (3): 387–416. Bibcode:2020JPal...94..387G. doi:10.1017/jpa.2020.1. S2CID 216505978.

[PABAraucaria-84] Robert Noll; Lutz Kunzmann (2020). "Diversity in fossil Araucaria Juss.: new species from the Middle Jurassic Jaramillo Petrified Forests in Santa Cruz province, Argentina". Palaeontographica Abteilung B. 301 (1–3): 3–75. Bibcode:2020PalAB.301....3N. doi:10.1127/palb/2020/0070. S2CID 224978809.

[85] Ruth A. Stockey; Gar W. Rothwell (2020). "Diversification of crown group Araucaria: the role of Araucaria famii sp. nov. in the mid-Cretaceous (Campanian) radiation of Araucariaceae in the Northern Hemisphere". American Journal of Botany. 107 (7): 1072–1093. doi:10.1002/ajb2.1505. PMID 32705687. S2CID 225568264.

[86] Gongle Shi; Haomin Li; Andrew B. Leslie; Zhiyan Zhou (2020). "Araucaria bract-scale complex and associated foliage from the early-middle Eocene of Antarctica and their implications for Gondwanan biogeography". Historical Biology: An International Journal of Paleobiology. 32 (2): 164–173. Bibcode:2020HBio...32..164S. doi:10.1080/08912963.2018.1472255. S2CID 89662557.

[87] Gabriella Rossetto-Harris; Peter Wilf; Ignacio H. Escapa; Ana Andruchow-Colombo (2020). "Eocene Araucaria Sect. Eutacta from Patagonia and floristic turnover during the initial isolation of South America". American Journal of Botany. 107 (5): 806–832. doi:10.1002/ajb2.1467. PMID 32388874. S2CID 218582967.

[88] L.C.A. Martínez; E. Pacheco Huacallo; R.R. Pujana; H. Padula (2020). "A new megaflora (leaves and reproductive structures) from the Huancané Formation (Lower Cretaceous), Peru". Cretaceous Research. 110: Article number 104426. Bibcode:2020CrRes.11004426M. doi:10.1016/j.cretres.2020.104426. S2CID 213340202.

[89] Zikun Jiang; Hao Wu; Ning Tian; Yongdong Wang; Aowei Xie (2020). "A new species of conifer wood Brachyoxylon from the Cretaceous of Eastern China and its paleoclimate significance". Historical Biology: An International Journal of Paleobiology. 33 (10): 1989–1995. doi:10.1080/08912963.2020.1755282. S2CID 219088846.

[90] Maria Edenilce P. Batista; Lutz Kunzman; Artur A. A. Sá; Antônio Álamo F. Saraiva; Maria Iracema B. Loiola (2020). "A new species of Brachyphyllum from the Crato Formation (Lower Cretaceous), Araripe Basin, Brazil". Ameghiniana. 57 (6): 519–533. doi:10.5710/AMGH.23.06.2020.3333. S2CID 226545919.

[91] Jiří Kvaček; Mário Miguel Mendes (2020). "Callialastrobus sousai gen. et sp. nov., a new araucariaceous pollen cone from the Early Cretaceous of Catefica (Lusitanian Basin, western Portugal) bearing Callialasporites and Araucariacites pollen". Review of Palaeobotany and Palynology. 283: Article 104313. Bibcode:2020RPaPa.28304313K. doi:10.1016/j.revpalbo.2020.104313. S2CID 225290387.

[92] Ünal Akkemik (2020). "A new fossil Cedrus species from the early Miocene of northwestern Turkey and its possible affinities". Palaeoworld. 30 (4): 746–756. doi:10.1016/j.palwor.2020.12.003. S2CID 230541679.

[93] Guoqing Xia; Ning Tian; Marc Philippe; Haisheng Yi; Chihua Wu; Gaojie Li; Zhiqiang Shi (2020). "Oldest Jurassic wood with Gondwanan affinities from the Middle Jurassic of Tibetan Plateau and its paleoclimatological and paleoecological significance". Review of Palaeobotany and Palynology. 281: Article 104283. Bibcode:2020RPaPa.28104283X. doi:10.1016/j.revpalbo.2020.104283. S2CID 224930021.

[94] Ruth A. Stockey; Gar W. Rothwell; Brian A. Atkinson (2020). "Late Cretaceous diversification of cupressaceous conifers: a taiwanioid seed cone from the Eden Main, Vancouver Island, British Columbia, Canada". International Journal of Plant Sciences. 181 (5): 529–541. doi:10.1086/708383. S2CID 218924040.

[95] Nadezhda I. Blokhina (2020). "Cupressinoxylon klimovii Blokhina, nom. nov. ‒ a new species name replacing Cupressinoxylon biotoides Blokhina, 1989 (Cupressaceae)". Botanica Pacifica. 9 (2): 196. doi:10.17581/bp.2020.09218.

[96] César Ríos-Santos; Sergio R.S. Cevallos-Ferriz; R.R. Pujana (2020). "Cupressaceous woods in the Upper Cretaceous Cabullona Group in Fronteras, Sonora, Mexico". Journal of South American Earth Sciences. 104: Article 102756. Bibcode:2020JSAES.10402756R. doi:10.1016/j.jsames.2020.102756. S2CID 224884535.

[97] Xu-Dong Gou; Sui Wan; Fu-Guang Zhao; Xin-Shi Cheng; Hai-Bo Wei; Yun Guo; Shi-Ling Yang; Zhuo Feng (2020). "A new conifer stem, Ductoagathoxylon wangii from the Middle Jurassic of the Santanghu Basin, Xinjiang, Northwest China". Review of Palaeobotany and Palynology. 285: Article 104357. doi:10.1016/j.revpalbo.2020.104357.

[98] Mário Miguel Mendes; Jiří Kvaček (2020). "Friisia lusitanica gen. et sp. nov., a new podocarpaceous ovuliferous cone from the Lower Cretaceous of Lusitanian Basin, western Portugal". Cretaceous Research. 108: Article 104352. Bibcode:2020CrRes.10804352M. doi:10.1016/j.cretres.2019.104352. S2CID 212921077.

[99] Ünal Akkemik (2020). "A new species of Juniperoxylon from the early Miocene of northwestern Turkey". Acta Palaeontologica Romaniae. 17 (1): 15–26. doi:10.35463/j.apr.2021.01.02.

[100] Ünal Akkemik; Dimitra Mantzouka; Demet Kıran Yıldırım (2020). "The first report of Lesbosoxylon Süss & Velitzelos from the early-middle Miocene of eastern Anatolia". Geodiversitas. 42 (23): 427–441. doi:10.5252/geodiversitas2020v42a23. S2CID 222210783.

[101] Andrey O. Frolov; Irina M. Mashchuk (2020). "Discovery of isolated leaves of Marskea (Taxaceae) in the Middle Jurassic sediments of Irkutsk Basin (East Siberia, Russia)". Phytotaxa. 449 (2): 164–172. doi:10.11646/phytotaxa.449.2.4. S2CID 225702863.

[102] Ruth A. Stockey; Harufumi Nishida; Gar W. Rothwell (2020). "Evolutionary diversification of taiwanioid conifers: evidence from a new Upper Cretaceous seed cone from Hokkaido, Japan". Journal of Plant Research. 133 (5): 681–692. Bibcode:2020JPlR..133..681S. doi:10.1007/s10265-020-01214-y. PMC 7429551. PMID 32686035.

[PalaeoworldZhouetal-103] Ling-Qi Zhou; Cuo Peng; Peng Deng; Xiao-Qin Zhang; Guo-Lin Yang; Wen-Xiu Ren; Jun Wang; Xiao-Qiang Li; Shi-Bo Tuo; Bing Guo (2020). "New records of Early Cretaceous petrified wood in Yumen, northwestern Gansu Province, China and their palaeoclimatic implications". Palaeoworld. 30 (3): 503–514. doi:10.1016/j.palwor.2020.08.002.

[104] Martina Dolezych; Lutz Reinhardt (2020). "First evidence for the conifer Pinus, as Pinuxylon selmeierianum sp. nov., during the Paleogene on Wootton Peninsula, northern Ellesmere Island, Nunavut, Canada". Canadian Journal of Earth Sciences. 57 (1): 25–39. Bibcode:2020CaJES..57...25D. doi:10.1139/cjes-2018-0163.

[105] Meng-Xiao Wu; Jian Huang; Tao Su; Qin Leng; Zhe-Kun Zhou (2020). "Tsuga seed cones from the late Paleogene of southwestern China and their biogeographical and paleoenvironmental implications". Palaeoworld. 29 (3): 617–628. doi:10.1016/j.palwor.2019.07.005. S2CID 199885815.

[106] Xiao Shi; Jianxin Yu; Yuewu Sun (2020). "Tyloses in the Lopingian cordaitalean root from Xinjiang, Northwest China". Review of Palaeobotany and Palynology. 273: Article 104134. Bibcode:2020RPaPa.27304134S. doi:10.1016/j.revpalbo.2019.104134. S2CID 210278581.

[107] Josefina Bodnar; Eduardo M. Morel; Eliana P. Coturel; Daniel G. Ganuza (2020). "New plant fossil records and biostratigraphic analysis from the Uspallata Group (Late Triassic) at Cacheuta Hill, Cuyo Basin, west-central Argentina". Geobios. 60: 3–27. Bibcode:2020Geobi..60....3B. doi:10.1016/j.geobios.2020.04.002. S2CID 219762016.

[108] Leyla J. Seyfullah; Emily A. Roberts; Alexander R. Schmidt; Eugenio Ragazzi; Ken B. Anderson; Daniel Rodrigues do Nascimento Jr.; Wellington Ferreira da Silva Filho; Lutz Kunzmann (2020). "Revealing the diversity of amber source plants from the Early Cretaceous Crato Formation, Brazil". BMC Evolutionary Biology. 20 (1): 107. Bibcode:2020BMCEE..20..107S. doi:10.1186/s12862-020-01651-2. PMC 7439571. PMID 32819273. S2CID 221183260.

[109] Serge V. Naugolnykh (2020). "Archaeopetalanthus progressus gen. et sp. nov. – a new representative of the vojnovskyopsid gymnosperms from the Carboniferous of Siberia (Russia)". Wulfenia. 27: 97–113.

[110] Mário Miguel Mendes; Kaj Raunsgaard Pedersen; Else Marie Friis (2020). "Battenispermum hirsutum gen. et sp. nov., a new Early Cretaceous seed from Portugal with chlamydospermous organisation". Cretaceous Research. 109: Article 104376. Bibcode:2020CrRes.10904376M. doi:10.1016/j.cretres.2020.104376. S2CID 213898064.

[111] María A. Gómez; Gabriela G. Puebla; Mercedes B. Prámparo; Andrea B. Arcucci (2020). "Fossil seeds from the La Cantera Formation, Early Cretaceous, San Luis Province, Argentina". Acta Palaeobotanica. 60 (1): 181–198. doi:10.35535/acpa-2020-0008. hdl:11336/145016.

[Ductolobatopitys-112] Domingas Maria da Conceição; Luiz Saturnino de Andrade; Rodrigo Neregato; Roberto Iannuzzi; Alexandra Crisafulli; Juan Carlos Cisneros (2020). "New petrified gymnosperms from the Permian of Maranhão (Pedra de Fogo Formation), Brazil: Ductolobatopitys nov. gen. and Kaokoxylon". Geobios. 60: 47–59. Bibcode:2020Geobi..60...47C. doi:10.1016/j.geobios.2020.04.003. S2CID 219737764.

[113] Wei-Ming Zhou; Bi-Yun Chen; Wei Sun; Xue-Zhi He; Jason Hilton; Jun Wang (2020). "A new gigantopterid genus from the late Permian of the Daha Coalfield, Tibetan Plateau and its implication on plant-insect interactions". Historical Biology: An International Journal of Paleobiology. 33 (12): 3228–3240. doi:10.1080/08912963.2020.1860033. S2CID 234390424.

[114] Sheng-Hui Deng; Xiao-Ju Yang; Zhi-Yan Zhou (2020). "A new Ginkgo from the Lower Cretaceous of Liaoning, Northeast China and its evolutionary implications". Review of Palaeobotany and Palynology. 283: Article 104315. Bibcode:2020RPaPa.28304315D. doi:10.1016/j.revpalbo.2020.104315. S2CID 225031049.

[115] Yong Yang; Yingwei Wang; David Kay Ferguson (2020). "A new macrofossil ephedroid plant with unusual bract morphology from the Lower Cretaceous Jiufotang Formation of northeastern China". BMC Evolutionary Biology. 20 (1): Article number 19. Bibcode:2020BMCEE..20...19Y. doi:10.1186/s12862-019-1569-y. PMC 7001366. PMID 32019502.

[Johniphyllum-116] Cindy V. Looy; Ivo A. P. Duijnstee (2020). "Voltzian conifers of the South Ash Pasture flora (Guadalupian, Texas): Johniphyllum multinerve gen. et sp. nov., Pseudovoltzia sapflorensis sp. nov., and Wantus acaulis gen. et sp. nov". International Journal of Plant Sciences. 181 (3): 363–385. doi:10.1086/706853. S2CID 208592093.

[117] Heidi M. Anderson; Maria Barbacka; Marion K. Bamford; W. B. Keith Holmes; John M. Anderson (2020). "Dicroidium (foliage) and affiliated wood Part 3 of a reassessment of Gondwana Triassic plant genera and a reclassification of some previously attributed". Alcheringa: An Australasian Journal of Palaeontology. 44 (1): 64–92. Bibcode:2020Alch...44...64A. doi:10.1080/03115518.2019.1622779. S2CID 199109037.

[118] Jana Čepičková; Jiří Kvaček (2020). "Two cycads Nilssonia mirovanae sp. nov. and Pseudoctenis babinensis J.Kvaček from the Cenomanian of the Bohemian Cretaceous Basin (the Czech Republic) as indicators of water stress in the palaeoenvironment". Fossil Imprint. 76 (2): 315–324. doi:10.37520/fi.2020.025.

[Novaiorquepitys-119] Domingas Maria da Conceição; Alexandra Crisafulli; Roberto Iannuzzi; Rodrigo Neregato; Juan Carlos Cisneros; Luiz Saturnino de Andrade (2020). "New petrified gymnosperms from the Permian of Maranhão (Pedra de Fogo Formation), Brazil: Novaiorquepitys and Yvyrapitys". Review of Palaeobotany and Palynology. 276: Article 104177. Bibcode:2020RPaPa.27604177D. doi:10.1016/j.revpalbo.2020.104177. S2CID 212954578.

[120] Sergey V. Naugolnykh; Valeryi V. Linkevich (2020). "Artinskian flora (Lower Permian) of the stratotypic area (the Middle Cis-Urals)". Socialno-ecologicheskie Technologii. 10 (2): 133–150. doi:10.31862/2500-2961-2020-10-2-133-150.

[121] Ming-Li Wan; Wan Yang; Jun Wang (2020). "Palaeocupressinoxylon uniseriale n. gen. n. sp., a gymnospermous wood from the upper Permian of Central Taodonggou, southern Bogda Mountains, northwestern China". Palaeoworld. 29 (1): 117–125. doi:10.1016/j.palwor.2019.06.002.

[122] Anastasia Zolina; Lina Golovneva; Natalya Nosova; Alexander Grabovskiy (2020). "A new species of Phoenicopsis (Leptostrobales) from the Maastrichtian–Danian of Chukotka, Russia". Geobios. 63: 67–75. Bibcode:2020Geobi..63...67Z. doi:10.1016/j.geobios.2020.09.002. S2CID 228928283.

[123] John G. Conran; Jennifer M. Bannister; Uwe Kaulfuss; Daphne E. Lee (2020). "Pterostoma neehoffii (cf. Zamiaceae): a new species of extinct cycad from the middle Miocene of New Zealand and an overview of fossil New Zealand cycads". New Zealand Journal of Botany. 58 (1): 30–47. Bibcode:2020NZJB...58...30C. doi:10.1080/0028825X.2019.1653939. S2CID 202847120.

[RPPDicroidium-124] Patrick Blomenkemper; Hans Kerp; Abdalla Abu Hamad; Benjamin Bomfleur (2020). "Contributions towards whole-plant reconstructions of Dicroidium plants (Umkomasiaceae) from the Permian of Jordan". Review of Palaeobotany and Palynology. 278: Article 104210. Bibcode:2020RPaPa.27804210B. doi:10.1016/j.revpalbo.2020.104210. S2CID 216248477.

[125] Yifan Hua; Xuelian Wang; Junlin Dong; Yanzhao Ji; Bainian Sun (2020). "A number of new seed fossils from the lower Permian of Gansu, Northwest China: implication for research on arils". Historical Biology: An International Journal of Paleobiology. 32 (8): 1098–1107. Bibcode:2020HBio...32.1098H. doi:10.1080/08912963.2019.1566323. S2CID 91993795.

[RPPUmaltolepis-126] Natalya Nosova (2020). "Female reproductive structures of Umaltolepis Krassilov and associated short shoots, buds and leaves of Pseudotorellia Florin from the Middle Jurassic of Angren, Uzbekistan". Review of Palaeobotany and Palynology. 281: Article 104266. Bibcode:2020RPaPa.28104266N. doi:10.1016/j.revpalbo.2020.104266. S2CID 219437430.

[127] Josef Pšenička; Jun Wang; Jason Hilton; Weiming Zhou; Jiří Bek; Stanislav Opluštil; Jana Votočková Frojdová (2020). "A small heterophyllous vine climbing on Psaronius and Cordaites trees in the earliest Permian forests of North China" (PDF). International Journal of Plant Sciences. 181 (6): 616–645. doi:10.1086/708814. S2CID 219926230.

[128] Weiming Zhou; Shan Wan; Mingli Wan; Jason Hilton; Josef Pšenička; Jun Wang (2020). "Yangopteris ascendens (Halle) gen. et comb. nov., a climbing alethopterid pteridosperm from the Asselian (earliest Permian) Wuda Tuff Flora". Review of Palaeobotany and Palynology. 294: Article 104282. doi:10.1016/j.revpalbo.2020.104282. S2CID 225462114.

[129] Pedro Correia; Arden R. Bashforth; Zbynĕk Šimůnek; Christopher J. Cleal; Artur A. Sá; Conrad C. Labandeira (2020). "The history of herbivory on sphenophytes: a new calamitalean with an insect gall from the Upper Pennsylvanian of Portugal and a review of arthropod herbivory on an ancient lineage". International Journal of Plant Sciences. 181 (4): 387–418. doi:10.1086/707105. S2CID 214292984.

[130] Ruiyun Li; Xiaoqiang Li; Xuelian Wang; Bainian Sun (2020). "First fossil liverwort with in situ flask-shaped receptacles from the Lower Cretaceous of Inner Mongolia, China". Cretaceous Research. 119: Article 104684. doi:10.1016/j.cretres.2020.104684. ISSN 0195-6671. S2CID 226314562.

[131] Xiao-Yuan He; Shi-Jun Wang; Jason Hilton; Jean Galtier; Hong-Guan Jiang (2020). "An advanced species of the fern Botryopteris Renault from the Permian of southwestern China" (PDF). Review of Palaeobotany and Palynology. 273: Article 104136. Bibcode:2020RPaPa.27304136H. doi:10.1016/j.revpalbo.2019.104136. S2CID 210266014.

[132] L. B. Golovneva; A. A. Grabovskiy; A. A. Zolina (2020). "A new species of the genus Birisia (Dicksoniaceae) from the lower–middle Albian deposits of Southern Primorye, Far East of Russia". Palaeobotany. 11: 74–95. doi:10.31111/palaeobotany/2020.11.74. S2CID 245112234.

[133] Barry A. Thomas (2020). "A new species of leafy calamite stem from the Pennsylvanian (Bolsovian) of the South Wales Coalfield". Acta Palaeobotanica. 60 (1): 207–211. doi:10.35535/acpa-2020-0010. hdl:2160/cd78c623-2a50-4c65-a758-7207027deacd.

[Catenuporella-134] Yueyang Zhang; Hongxia Jiang; Yasheng Wu; Hongping Bao; Junfeng Ren; Zhengliang Huang (2020). "Calcified dasycladaleans from the Upper Ordovician in the Ordos Basin, China". Acta Micropalaeontologica Sinica. 37 (3): 228–237. doi:10.16087/j.cnki.1000-0674.2020.03.003.

[auto-135] Jordi Pérez-Cano; Telm Bover-Arnal; Carles Martín-Closas (2020). "Barremian charophytes from the Maestrat Basin". Cretaceous Research. 115. doi:10.1016/j.cretres.2020.104544. hdl:2445/171586. S2CID 224875084.

[136] Jan Hinkelman; Lucia Vršanská (2020). "A Myanmar amber cockroach with protruding feces contains pollen and a rich microcenosis". The Science of Nature. 107 (2): Article number 13. Bibcode:2020SciNa.107...13H. doi:10.1007/s00114-020-1669-y. PMID 32125545. S2CID 211730431.

[CR109Santamarinaetal-137] Patricio Emmanuel Santamarina; Viviana Dora Barreda; Ari Iglesias; Augusto Nicolás Varela (2020). "Palynology from the Cenomanian Mata Amarilla Formation, southern Patagonia, Argentina". Cretaceous Research. 109: Article 104354. Bibcode:2020CrRes.10904354S. doi:10.1016/j.cretres.2019.104354. S2CID 212976442.

[BucureetalRIPS-138] ^ ^a ^b ^c ^d ^e ^f Ioan I. Bucur; Sylvain Rigaud; Nicolò Del Piero; Andrea Fucelli; Eric Heerwagen; Camille Peybernes; Giovan Peyrotty; Christian Verard; Jérôme Chablais; Rossana Martini (2020). "Upper Triassic calcareous algae from the Panthalassa Ocean". Rivista Italiana di Paleontologia e Stratigrafia. 126 (2): 499–540. doi:10.13130/2039-4942/13681.

[139] Xiu-Cai Yuan; Cong-Hui Xiong; Fan-Kai Sun; Zi-Xi Wang; Teng Mao; Yi-Jie Li; Chun-Hui Liu; Ming-Xuan Sun; Jun-Ling Dong; Bai-Nian Sun (2020). "The geological significance of a new species of Coniopteris from the Middle Jurassic of northwestern China". Historical Biology: An International Journal of Paleobiology. 32 (2): 267–280. Bibcode:2020HBio...32..267Y. doi:10.1080/08912963.2018.1488251. S2CID 89752285.

[140] Barry A. Thomas; Christopher J. Cleal (2020). "The nomenclature of fossil-taxa representing different preservational states: Lepidodendron as a case-study". Taxon. 69 (5): 1052–1061. doi:10.1002/tax.12291. S2CID 225235565.

[141] Yang Yu; San-Ping Xie; John Devaney; Si-Hang Zhang; Tian-Yu Chen; Xu Zeng; Bing Wang; Yu Zhang (2020). "A new species of Drynaria (Polypodiaceae) from the late Miocene of Yunnan, Southwest China and implications on the genus evolution" (PDF). Palaeobiodiversity and Palaeoenvironments. 100 (4): 939–949. Bibcode:2020PdPe..100..939Y. doi:10.1007/s12549-020-00429-0. S2CID 219730737.

[142] Li Zhang; Yong Wang; Hong-Yu Chen; Lei Han; Yu-Xin Zhang; Wen-Jia Li; Tao Yang; Hao-Jian Wang; Lin Bao; De-Fei Yan (2020). "New fossil material of Equicalastrobus (Equisetales) and associated leaves from the Late Triassic of Baojishan basin, Gansu Province, China". Historical Biology: An International Journal of Paleobiology. 33 (9): 1522–1533. doi:10.1080/08912963.2020.1716747. S2CID 213295920.

[PhytoKeys138-143] Aye Thida Aung; Jian Huang; Truong Van Do; Ai Song; Jia Liu; Zhe-Kun Zhou; Tao Su (2020). "Three new fossil records of Equisetum (Equisetaceae) from the Neogene of south-western China and northern Vietnam". PhytoKeys (138): 3–15. doi:10.3897/phytokeys.138.38674. PMC 6969027. PMID 31988601.

[144] Branko Sokač; Tonći Grgasović (2020). "New dasycladalean alga with unusual two types of laterals from the Palaocene deposits of Konavle, SE of Dubrovnik (Dinarides, Croatia)". Revue de Micropaléontologie. 69: Article 100464. Bibcode:2020RvMic..6900464S. doi:10.1016/j.revmic.2020.100464. S2CID 226343559.

[145] Ya Li; Yong-Dong Wang; Harald Schnerder; Peng-Cheng Wu (2020). "Frullania partita sp. nov. (Frullaniaceae, Porellales), a new leafy liverwort from the mid-Cretaceous of Myanmar". Cretaceous Research. 108: Article 104341. Bibcode:2020CrRes.10804341L. doi:10.1016/j.cretres.2019.104341. S2CID 213553976.

[146] Yuriy S. Mamontov; John J. Atwood; Evgeny E. Perkovsky; Michael S. Ignatov (2020). "Hepatics from Rovno amber (Ukraine): Frullania pycnoclada and a new species, F. vanae". The Bryologist. 123 (3): 421–430. doi:10.1639/0007-2745-123.3.421. S2CID 225406886.

[Gippslandites-147] Fearghus R. McSweeney; Jeff Shimeta; John St. J. S. Buckeridge (2020). "Two new genera of early Tracheophyta (Zosterophyllaceae) from the upper Silurian–Lower Devonian of Victoria, Australia". Alcheringa: An Australasian Journal of Palaeontology. 44 (3): 379–396. Bibcode:2020Alch...44..379M. doi:10.1080/03115518.2020.1744725. S2CID 218993880.

[148] Kelly C. Pfeiler; Alexandru M. F. Tomescu (2020). "An Early Devonian actinostelic euphyllophyte with secondary growth from the Emsian of Gaspé (Canada) and the importance of tracheid wall thickening patterns in early euphyllophyte systematics". Papers in Palaeontology. 7 (2): 1081–1095. doi:10.1002/spp2.1335. S2CID 225425418.

[149] Weiming Zhou; Josef Pšenička; Jiří Bek; Mingli Wan; C. Kevin Boyce; Jun Wang (2020). "A new anachoropterid fern from the Asselian (Cisuralian) Wuda Tuff Flora". Review of Palaeobotany and Palynology. 294: Article 104346. doi:10.1016/j.revpalbo.2020.104346. S2CID 228855095.

[150] Steven T. LoDuca; Anthony L. Swinehart; Matthew A. LeRoy; Denis K. Tetreault; Shawn Steckenfinger (2020). "Codium-like taxa from the Silurian of North America: morphology, taxonomy, paleoecology, and phylogenetic affinity". Journal of Paleontology. 95 (2): 207–235. doi:10.1017/jpa.2020.85. S2CID 228938523.

[151] Li-jun Chen; Ye-mao Hou; Peng-fei Yin; Xin Wang (2020). "An edible fruit from the Jurassic of China". China Geology. 3 (1): 8–15. Bibcode:2020CGeo....3....8C. doi:10.31035/cg2020010.

[152] Antoine Champreux; Brigitte Meyer-Berthaud; Anne-Laure Decombeix (2020). "Keraphyton gen. nov., a new Late Devonian fern-like plant from Australia". PeerJ. 8: e9321. doi:10.7717/peerj.9321. PMC 7304422. PMID 32587800.

[153] D. S. Kopylov; A. P. Rasnitsyn; D. S. Aristov; A. S. Bashkuev; N. V. Bazhenova; V. Yu. Dmitriev; A. V. Gorochov; M. S. Ignatov; V. D. Ivanov; A. V. Khramov; A. A. Legalov; E. D. Lukashevich; Yu. S. Mamontov; S. I. Melnitsky; B. Ogłaza; A. G. Ponomarenko; A. A. Prokin; O. V. Ryzhkova; A. S. Shmakov; N. D. Sinitshenkova; A. Yu. Solodovnikov; O. D. Strelnikova; I. D. Sukacheva; A. V. Uliakhin; D. V. Vasilenko; P. Wegierek; E. V. Yan; M. Zmarzły (2020). "The Khasurty Fossil Insect Lagerstätte". Paleontological Journal. 54 (11): 1221–1394. Bibcode:2020PalJ...54.1221K. doi:10.1134/S0031030120110027. S2CID 231850225.

[154] Yu.V. Mosseichik (2020). "Lepidodendron species from the Viséan of the Moscow Basin" (PDF). Lethaea Rossica. 20: 19–33.

[155] Yi-zhi Xu; Ning Yue; Shi Liu; Yi Zhang; Shao-lin Zheng; Yong-dong Wang; Xin-ran Huo (2020). "Lobatannularia linjiaensis sp. nov. from the Middle Triassic of Benxi, Liaoning". Acta Palaeontologica Sinica. 59 (3): 329–344. doi:10.19800/j.cnki.aps.2019.024.

[156] Sergey V. Naugolnykh; Ming Tu; Xiao-Yan Liu; Jian-Hua Jin (2020). "A new species of Lygodium (Schizaeaceae) from the Buxin Formation (Paleocene), Sanshui Basin, South China". Palaeoworld. 29 (3): 606–616. doi:10.1016/j.palwor.2019.07.003. S2CID 199101009.

[157] Tomoyuki Katagiri; Hisao Shinden (2020). "Discovery of a simple thalloid liverwort Metzgeriites kujiensis (Metzgeriaceae), a new species from Late Cretaceous Japanese amber". Hattoria. 11: 13–21. doi:10.18968/hattoria.11.0_13.

[158] Silvia Gnaedinger; Bárbara Cariglino; Ana María Zavattieri; Mariana Monti; Pedro R. Gutiérrez (2020). "Neoarthropitys gondwanaensis gen. et sp. nov. from the Middle Triassic of Gondwana: An intermediate stage in the anatomical trend of Equisetalean stems". Review of Palaeobotany and Palynology. 282: Article 104298. Bibcode:2020RPaPa.28204298G. doi:10.1016/j.revpalbo.2020.104298. S2CID 225347405.

[GFOsmundacaulis-159] Yeming Cheng; Fengxiang Liu; Xiaonan Yang; Tongxing Sun (2020). "Two new species of Mesozoic tree ferns (Osmundaceae: Osmundacaulis) in Eurasia as evidence of long-term geographic isolation". Geoscience Frontiers. 11 (5): 1875–1888. Bibcode:2020GeoFr..11.1875C. doi:10.1016/j.gsf.2020.01.019.

[AlcheringaZavattierietal-160] Ana M. Zavattieri; Pedro R. Gutiérrez; Mariana Monti (2020). "Middle Triassic freshwater green algae and fungi of the Puesto Viejo Basin, central-western Argentina: palaeoenvironmental implications". Alcheringa: An Australasian Journal of Palaeontology. 44 (3): 430–459. Bibcode:2020Alch...44..430Z. doi:10.1080/03115518.2020.1749302. S2CID 219519003.

[161] Li Liu; Josef Pšenička; Jiří Bek; Mingli Wan; Hermann W. Pfefferkorn; Jun Wang (2020). "A whole calamitacean plant Palaeostachya guanglongii from the Asselian (Permian) Taiyuan Formation in the Wuda Coalfield, Inner Mongolia, China". Review of Palaeobotany and Palynology. 294: Article 104245. doi:10.1016/j.revpalbo.2020.104245. S2CID 219017989.

[162] Ruiyun Li; Xiaoqiang Li; Shenghui Deng; Bainian Sun (2020). "Morphology and microstructure of Pellites hamiensis nov. sp., a Middle Jurassic liverwort from northwestern China and its evolutionary significance". Geobios. 62: 23–29. Bibcode:2020Geobi..62...23L. doi:10.1016/j.geobios.2020.07.003. S2CID 225500594.

[163] P. Hiller; M. Böhme; S. Schneider; J. Prieto; B. Bomfleur (2020). "Plenasium (Aurealcaulis) elegans sp. nov. from the Eocene of Vietnam – a connecting link in the evolution of modern Royal Ferns (Osmundeae, Osmundaceae)". Journal of Systematic Palaeontology. 18 (8): 703–715. Bibcode:2020JSPal..18..703H. doi:10.1080/14772019.2019.1683771. S2CID 209573114.

[164] Yi Zhang; Yong-Dong Wang; Yue Hong; Liu Cao; Fu-liang Gao (2020). "Pleuromeia discovered from the Middle Triassic Linjia Formation of Benxi, Northeast China". Palaeoworld. 29 (4): 706–714. doi:10.1016/j.palwor.2020.02.004. S2CID 216475501.

[165] Carmen Álvarez-Vázquez; Jiří Bek; Jana Drábková (2020). "Polysporia baetica sp. nov., a new heterosporous sub-arborescent isoetalean from lower Bolsovian (Middle Pennsylvanian) strata of the Peñarroya-Belmez-Espiel Coalfield (Córdoba, SW Spain)". Review of Palaeobotany and Palynology. 272: Article 104115. Bibcode:2020RPaPa.27204115A. doi:10.1016/j.revpalbo.2019.104115.

[166] Zlatko Kvaček; Vasilis Teodoridis (2020). "A new Oligocene fern of Dryopteridaceae from the Českéstředohoří Mts (Czech Republic)". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 295 (1): 9–16. doi:10.1127/njgpa/2020/0864. S2CID 213778635.

[167] Chunxiang Li; Robbin C. Moran; Junye Ma; Bo Wang; Jiasheng Hao (2020). "A new fossil record of Lindsaeaceae (Polypodiales) from the mid-Cretaceous amber of Myanmar". Cretaceous Research. 105: Article 104040. Bibcode:2020CrRes.10504040L. doi:10.1016/j.cretres.2018.12.010.

[168] Qing Tang; Ke Pang; Xunlai Yuan; Shuhai Xiao (2020). "A one-billion-year-old multicellular chlorophyte". Nature Ecology & Evolution. 4 (4): 543–549. Bibcode:2020NatEE...4..543T. doi:10.1038/s41559-020-1122-9. PMC 8668152. PMID 32094536. S2CID 211266015.

[charophytesGeobios-169] Josep Sanjuan; Ingeborg Soulié-Märsche (2020). "New charophyte assemblage from middle Miocene lacustrine deposits of Moneva (Ebro Basin, Spain)". Geobios. 59: 79–90. Bibcode:2020Geobi..59...79S. doi:10.1016/j.geobios.2020.03.001. S2CID 218775725.

[170] Pu Huang; Le Liu; Lu Liu; Min Qin; De-Ming Wang; Jin-Zhuang Xue (2020). "A new plant with novel leaves from the Upper Devonian of Zhejiang Province, China". Palaeoworld. 29 (4): 695–705. doi:10.1016/j.palwor.2020.03.003. S2CID 216265575.

[171] Mingli Wan; Wenjun Sun; Jiří Bek; Feng Liu; Christopher Hill; Jun Wang (2020). "Scolecopteris minuta sp. nov., a marattialean fern from the early Permian Wuda Tuff Flora of Inner Mongolia, China". Review of Palaeobotany and Palynology. 294: Article 104246. doi:10.1016/j.revpalbo.2020.104246. S2CID 219013492.

[RPPSigillaria-172] Michael P. D'Antonio; C. Kevin Boyce; Jun Wang (2020). "Two new species of Sigillaria Brongniart from the Wuda Tuff (Asselian: Inner Mongolia, China) and their implications for lepidodendrid life history reconstruction". Review of Palaeobotany and Palynology. 294: Article 104203. doi:10.1016/j.revpalbo.2020.104203. S2CID 216296882.

[173] Chunxiang Li; Robbin C. Moran; Junye Ma; Bo Wang; Jiasheng Hao; Qun Yang (2020). "A mid-Cretaceous tree fern of Thyrsopteridaceae (Cyatheales) preserved in Myanmar amber". Cretaceous Research. 105: Article 104050. Bibcode:2020CrRes.10504050L. doi:10.1016/j.cretres.2019.01.002.

[174] Zhuo Feng; Hai-Bo Wei; Yun Guo; Xiao-Yuan He; Qun Sui; Yu Zhou; Hang-Yu Liu; Xu-Dong Gou; Yong Lv (2020). "From rainforest to herbland: New insights into land plant responses to the end-Permian mass extinction". Earth-Science Reviews. 204: Article 103153. Bibcode:2020ESRv..20403153F. doi:10.1016/j.earscirev.2020.103153.

[175] Serge V. Naugolnykh (2020). "A new species of the genus Tumidopteris Naugolnykh from the Permian of the Pechora Cis-Urals, Russia". Fossil Imprint. 76 (2): 270–278. doi:10.37520/fi.2020.022.

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