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Peopling of Southeast Asia

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(Redirected from Genetic history of Thailand)
See Archaic humans in Southeast Asia for the earlier presence of archaic humans.

Southeast Asia was first reached by anatomically modern humans possibly before 70,000 years ago.[1] Anatomically modern humans are suggested to have reached Southeast Asia twice in the course of the Southern Dispersal migrations during and after the formation of a distinct East Asian clade from 70,000 to 50,000 years ago.[2][3]

Archeological discoveries

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In Asia, the most recent late archaic human fossils were found in Thailand (125-100 ka), the Philippines (58-24 ka), Malaysia (c. 40 ka), and Sri Lanka (c.36 ka).[4] The artifacts from these sites include partial skeleton, crania, deep skull, and other related skeletons indicate that modern human migrated to Asia earlier than the western theory might have discussed.[5]

In 2007, an analysis of cut marks on two bovid bones found in Sangiran, showed them to have been made 1.5 to 1.6 million years ago by clamshell tools. This may be the oldest evidence for the presence of early humans in today Indonesia and are to date the oldest evidence of shell tool use in the world.[6]

In 2009, archaeologists discovered the partial cranium and some teeth of a modern human at Tam Pa Ling in mainland Laos, which shed light on the understanding of anatomically modern human migration and evolution in the region during the Late Pleistocene Period.[5] The site is located in Houaphanh Province, around 170 miles north of Vientiane, the capital city of modern Laos. Within this site, only human remains were found, and there is no evidence of human occupation or other artifacts. The radiocarbon dating of the charcoal and the sediment dating analyses identify the remains to date at least c. 56.5 ka, while the dental artifacts from the remains that analyzed by the isotope-ratio measurement indicate c. 63.6 ka.[5] The analysis of the cranium and dentition of the remains suggest that these are the remains of early modern human populations in Southeast Asia. This date is older than the fossils that were found in Niah cave in Malaysia, which offers another explanation for human evolution in Southeast Asia.

In addition to the discovery in Laos, there are also a number of human remains and related artifacts found across mainland Southeast Asia in which it suggests the new ideas of the regional Late Pleistocene development as well. More teeth and molars that were found in Thailand and Vietnam sites (Tham Wihan Naki, Thailand; Tham Kuyean, Vietnam, etc.) indicate transitions between H. erectus and H. sapiens.[7] In fact, these remains might indicate the possible interbreeding between H. erectus and H. sapiens, such as the tooth at Wihan Nakin at Chaiyaphum province in Thailand.[7]

Migrations

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The earliest modern human inhabitants of Southeast Asia were hunter-gatherers that arrived in the area at least c. 40,000 BP. Contemporary remnant groups of these earliest inhabitants (e.g. the Semang of Malaysia or the Aetas of the Philippines) are usually included under the cover term "Negrito".[8] The earliest settlers had sufficient maritime technology to cross the Wallace Line, probably at a similar date to the first settlement of Sahul (c. 45,000 BP/49,000 – 43,000 BP).[9]: 50 

The Neolithic was characterized by several migrations from southern China into Mainland and Island Southeast Asia by Austroasiatic and Austronesian-speakers.[8][10]

The proposed route of Austroasiatic and Austronesian migration into Insular Southeast Asia during the Neolithic period. (Simanjuntak, 2017)[11]

During the Neolithic, Austroasiatic peoples populated Indochina via land routes. The earliest agricultural societies that cultivated millet and wet-rice emerged around 1700 BCE in the lowlands and river floodplains of Indochina.[12] Based on archaeological and genetic evidence, it is assumed that Austroasiatic speakers also expanded into Insular Southeast Asia in the Neolithic, but were later supplanted or assimilated by Austronesian speakers.[11]

The Austronesian Expansion(3500 BC to AD 1200)

The most widespread migration event was the Austronesian expansion, which began at around 5,500 BP (3500 BC) from coastal southern China via Taiwan. Due to their use of ocean-going outrigger boats and voyaging catamarans,[a] Austronesians rapidly colonized Island Southeast Asia, before spreading further into Micronesia, Melanesia, Polynesia, Madagascar[b] and the Comoros. They dominated the lowlands and coasts of Island Southeast Asia, giving rise to modern Islander Southeast Asians, Micronesians, Polynesians, and Malagasy.[14][15][16][17] The first Austronesians reached the Philippines at around 2200 BC, settling the Batanes Islands and northern Luzon from Taiwan. From there, they rapidly spread downwards to the rest of the islands of the Philippines and Southeast Asia.[18][19]

The widespread presence of Kra-Dai, Tibeto-Burman, and Hmong-Mien speakers in Mainland Southeast Asia is the result of later migrations. Originating from southern China, where many languages of these families are still spoken, they expanded southwards into Southeast Asia in historical times around the second half of the first millennium CE.[8]

Trade

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Territorial principalities in both Insular and Mainland Southeast Asia, characterised as Agrarian kingdoms[20] had by around 500 BCE developed an economy based on surplus crop cultivation and moderate coastal trade of domestic natural products. Several states of the Malayan-Indonesian "thalassian" zone[21] shared these characteristics with Indochinese polities like the Pyu city-states in the Irrawaddy river valley, Van Lang in the Red River delta and Funan around the lower Mekong.[22] Văn Lang, founded in the 7th century BCE endured until 258 BCE under the rule of the Hồng Bàng dynasty, as part of the Đông Sơn culture eventually sustained a dense and organised population, that produced an elaborate Bronze Age industry.[23][24]

Intensive wet-rice cultivation in an ideal climate enabled the farming communities to produce a regular crop surplus, that was used by the ruling elite to raise, command and pay work forces for public construction and maintenance projects such as canals and fortifications.[23][21] Though millet and rice cultivation was introduced around 2000 BCE, hunting and gathering remained an important aspect of food provision, in particular in forested and mountainous inland areas.[25]

Historians have emphasized the maritime connectivity of the Southeast Asian region whereby it can be analyzed as a single cultural and economic unit, as has been done with the Mediterranean basin.[26] This region stretches from the Yangtze delta in China down to the Malay Peninsula, including the South China Sea, Gulf of Thailand and Java Sea. The region was dominated by the thalassocratic cultures of the Austronesian peoples.[27][28][29]

Genetics

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See also: Hoabinhian ancestry and Ancient Southern East Asian
Estimated ancestry components among selected modern populations per Changmai et al (2022). The yellow component represents East Asian-like ancestry.[30]
Principal component analysis (PCA) of ancient and present-day individuals from worldwide populations after the out-of-Africa expansion.

One study (Chaubey 2015) found evidence for ancient gene flow from East Asian-related groups into the Andamanese people, suggesting that Andamanese (Onge) had about 30% East Asian-related ancestry next to their original Negrito ancestry, though the authors also suggest that this latter finding may in fact reflect the genetic affinity of the Andamanese to Melanesian, Southeast Asian, and Asian Negrito populations rather than true East Asian admixture (stating that "The Han ancestry measured in Andaman Negrito is probably partially capturing both Melanesian and Malaysian Negrito ancestry"),[31] as a previous study by the authors (Chaubey et al.) indicated "a deep common ancestry" between Andamanese, Melanesians and other Negrito groups (as well as South Asians), and an affinity between Southeast Asian Negritos and Melanesians with East Asians.[32]

According to McColl et. al (2018), present-day Southeast Asians derive their ancestry from at least four ancient populations. The oldest source derives from mainland Hòabìnhians, who share ancestry with present-day Andamanese Önge, Malaysian Jehai, and the ancient Japanese Ikawazu Jōmon. During the Neolithic period, East Asian farmers intermixed with the native inhabitants and contrary to popular opinion, did not replace them. These farmers also shared ancestry with present-day Austroasiatic-speaking hill tribes themselves. About 2000 years ago, new East Asian ancestry components were introduced and coincided with the introduction of ancestral Kradai languages and Austronesian culture in mainland and maritime Southeast Asia respectively.[33]

A 2020 genetic study on Southeast Asian populations focusing on ethnic groups in Vietnam by Liu et al. 2020 found that most sampled groups are closely related to East Asians and carry mostly "East Asian-related" ancestry. Modern Austronesian and Austroasiatic speaking populations of Southeast Asia were found to have mostly East Asian-related ancestry (89% to 96%, with 94% on average). Taiwanese indigenous peoples had on average 99% East Asian-related ancestry. Kra–Dai-speaking populations had, similar to the Taiwanese indigenous peoples, nearly exclusively East Asian-related ancestry.[34]

A recent study from 2021 found that an ancient Holocene hunter-gatherer from South Sulawesi had ancestry from both a distinct lineage related to modern Papuans and Aboriginal Australians and from the East-Eurasian lineage (represented by modern East Asians). The hunter-gatherer individual had approximately ~50% "Basal-East Asian" ancestry, and was positioned in between modern East Asians and Papuans of Oceania. The authors concluded that East Asian-related ancestry expanded much earlier into Maritime Southeast Asia than previously suggested, long before the expansion of Austroasiatic and Austronesian groups.[35]

Another study about the ancestral composition of modern ethnic groups in the Philippines from 2021 suggests that distinctive Basal-East Asian (East-Eurasian) ancestry originated in Mainland Southeast Asia at ~50,000BC, and expanded through multiple migration waves southwards and northwards respectively.[3]

A 2022 study stated that there was substantial South Asian admixture in various Southeast Asian populations in Thailand, Cambodia, Vietnam, Myanmar and Singapore. Exceptions were isolated hill tribes and present hunter-gatherer groups in Thailand. Admixture rates for Southeast Asians range from 2% to 16% and originated from Indian cultural influence in the region.[36]

See also

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Notes

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  1. ^ Different dates are argued for the introduction of the various pieces of maritime technology by Austronesians. These are based on linguistics and the distribution of types on first European encounter. There is no early archaeological evidence, and little iconographic or written evidence on this until the first encounters with Europeans
  2. ^ c.7th century CE or later[13]

References

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  2. ^ Lipson, Mark; Reich, David (2017). "A Working Model of the Deep Relationships of Diverse Modern Human Genetic Lineages Outside of Africa". Molecular Biology and Evolution. 34 (4): 889–902. doi:10.1093/molbev/msw293. PMC 5400393. PMID 28074030. The former [eastern clade] includes present-day East Asians and had differentiated as early as the ~40 kya Tianyuan individual (Fu et al. 2013), while early members of the latter [western clade] include ancient European hunter-gatherers (Lazaridis et al. 2014; Seguin-Orlando et al. 2014; Fu et al. 2016) and the ancient northern Eurasian Mal'ta 1 (MA1, a ~24 kya Upper Paleolithic individual from south-central Siberia) (Raghavan et al. 2014). More recent (Neolithic and later) western Eurasians, such as Europeans, are mostly descended from the western clade but with an additional component of "Basal Eurasian" ancestry (via the Near East) splitting more deeply than any other known non-African lineage (Lazaridis et al. 2014, 2016). The timing of the eastern/western split is uncertain, but several papers (Gutenkunst et al. 2009; Laval et al. 2010; Gravel et al. 2011) have used present-day European and East Asian populations to infer dates of initial separation of 40–45 kya (adjusted for a mutation rate of 0.5 × 10−9 per year; Scally 2016).
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  35. ^ Carlhoff, Selina; Duli, Akin; Nägele, Kathrin; Nur, Muhammad; Skov, Laurits; Sumantri, Iwan; Oktaviana, Adhi Agus; Hakim, Budianto; Burhan, Basran; Syahdar, Fardi Ali; McGahan, David P. (August 2021). "Genome of a middle Holocene hunter-gatherer from Wallacea". Nature. 596 (7873): 543–547. Bibcode:2021Natur.596..543C. doi:10.1038/s41586-021-03823-6. hdl:10072/407535. ISSN 1476-4687. PMC 8387238. PMID 34433944. The qpGraph analysis confirmed this branching pattern, with the Leang Panninge individual branching off from the Near Oceanian clade after the Denisovan gene flow, although with the most supported topology indicating around 50% of a basal East Asian component contributing to the Leang Panninge genome (Fig. 3c, Supplementary Figs. 7–11).
  36. ^ Changmai, Piya; Jaisamut, Kitipong; Kampuansai, Jatupol; et al. (2022). "Indian genetic heritage in Southeast Asian populations". PLOS Genetics. 18 (2): e1010036. doi:10.1371/journal.pgen.1010036. PMC 8853555. PMID 35176016.

Further reading

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