Pygopodoidea

Pygopodoidea; Temporal range: Albian – Present, 110–0 Ma PreꞒ Ꞓ O S D C P T J K Pg N
	Eastern stone gecko (Diplodactylus vittatus)
	Pink-tailed worm-lizard (Aprasia parapulchella)
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Order:	Squamata
Clade:	Gekkonomorpha
Infraorder:	Gekkota
Clade:	Pygopodomorpha; Vidal & Hedges, 2009
Superfamily:	Pygopodoidea; Gray, 1845
Families
	Diplodactylidae; Carphodactylidae; Pygopodidae;

Pygopodoidea is a gecko superfamily and the only taxon in the gekkotan subclade Pygopodomorpha. The clade includes three Australasian families: Diplodactylidae (stone geckos), Carphodactylidae (knob-tailed geckos), and Pygopodidae (flap-footed geckos). Traditional gekkotan systematics had considered Diplodactylidae and Carphodactylidae as subfamilies of the family Gekkonidae, but recent molecular work have placed Pygopodidae within Gekkonidae making it paraphyletic.^[1] These analyses have shown support of Pygopodidae and Carphodactylidae being sister taxa, with Diplodactylidae occupying a basal position in Pygopodoidea.^[2]^[3]^[4]

Below is a taxonomic list of pygopodoid genera in taxonomic order:

Pygopodomorpha (Vidal & Hedges, 2009)
- Pygopodoidea (Gray, 1845)
  - Diplodactylidae (Underwood, 1954) – Stone Geckos
    - Crenadactylus (Dixon & Kluge, 1964) – Clawless Gecko
    - Nebulifera (Oliver, Bauer, Greenbaum, Jackman & Hobbie, 2012) – Robust Velvet Gecko
    - Amalosia (Wells & Wellington, 1984) – Amalosian Velvet Geckos
    - Oedura (J. E. Gray, 1842) – Oeduran Velvet Geckos
    - Hesperoedura (Oliver, Bauer, Greenbaum, Jackman & Hobbie, 2012) – Reticulated Velvet Gecko
    - Strophurus (Fitzinger, 1843) – Spiny-tailed Geckos
    - Diplodactylus (Gray, 1832) – Stone Geckos
    - Rhynchoedura (Günther, 1867) – Beaked Geckos
    - Lucasium (Wermuth, 1965) – Ground Geckos
    - Toropuku (Nielsen, Bauer, Jackman, Hitchmough & Daugherty, 2011) – New Zealand Striped Gecko
    - Naultinus (Gray, 1842) – New Zealand Green Geckos
    - Tukutuku (Nielsen, Bauer, Jackman, Hitchmough & Daugherty, 2011) – Harlequin Gecko
    - Dactylocnemis (Steindachner, 1867) – Pacific Geckos
    - Mokopirirakau (Nielsen, Bauer, Jackman, Hitchmough & Daugherty, 2011) – Forest Geckos
    - Woodworthia (Garman, 1901) – Woodworth's Geckos
    - Hoplodactylus (Fitzinger, 1843) – Hoplodactyl Geckos
    - Pseudothecadactylus (Brongersma, 1936) – Cave Geckos
    - Bavayia (Roux, 1913) – Bavay's Geckos
    - Paniegekko (Bauer, Jackman, Sadlier & Whitaker, 2012) – Panié du Massif Gecko
    - Dierogekko (Bauer, Jackman, Sadlier & Whitaker, 2006) – New Caledonian Striped Geckos
    - Oedodera (Bauer, Jackman, Sadlier & Whitaker, 2006) – Marbled Gecko
    - Correlophus (Guichenot, 1866) – Correloph Geckos
    - Rhacodactylus (Fitzinger, 1843) – Rhacodactyl Geckos
    - Mniarogekko (Bauer, Whitaker, Sadlier & Jackman, 2012) – Mossy Geckos
    - Eurydactylodes (Wermuth, 1965) – New Caledonian Chameleon Geckos
  - Carphodactylidae (Kluge, 1967) – Knob-tailed Geckos
    - Phyllurus (Schinz, 1822) – Phyllur Leaf-tailed Geckos
    - Saltuarius (Couper, Covacevich & Moritz, 1993) – Saltuar Leaf-tailed Geckos
    - Orraya (Couper, Covacevich, Schneider & Hoskin, 2000) – McIlwraith Leaf-tailed Gecko
    - Carphodactylus (Günther, 1897) – Australian Chameleon Gecko
    - Uvidicolus (Oliver & Bauer, 2011) – Border Thick-tailed Gecko
    - Underwoodisaurus (Wermuth, 1965) – Underwood's Geckos
    - Nephrurus (Günther, 1876) – Knob-tailed Geckos
  - Pygopodidae (Gray, 1845) – Flap-footed Geckos
    - Delma (Gray, 1831) – Delmas
    - Lialis (Gray, 1835) – Lialis
    - Ophidiocephalus (Lucas & Frost, 1897) – Bronzeback Snake-lizard
    - Pletholax (Cope, 1864) – Slender Slider
    - Aprasia (Gray, 1839) – Worm Geckos
    - Paradelma (Kinghorn, 1926) – Brigalow Scalyfoot
    - Pygopus (Merrem, 1820) – Scalyfeet

References

^ Han, D., K. Zhou, & A. M. Bauer. 2004. Phylogenetic relationships among gekkotan lizards inferred from c-mos nuclear DNA sequences and a new classification of the Gekkota. Biological Journal of the Linnean Society 83: 353– 368.
^ Gamble, T., A. M. Bauer, E. Greenbaum, & T.R. Jackman. 2008. Evidence for Gondwanan vicariance in an ancient clade of gecko lizards. Journal of Biogeography 35: 88-104
^ Pyron, R; Burbrink, Frank T; Wiens, John J (2013). "A phylogeny and revised classification of Squamata, including 4161 species of lizards and snakes". BMC Evolutionary Biology. 13: 93. Bibcode:2013BMCEE..13...93P. doi:10.1186/1471-2148-13-93. PMC 3682911. PMID 23627680.
^ Zheng, Yuchi; Wiens, John J. (2016). "Combining phylogenomic and supermatrix approaches, and a time-calibrated phylogeny for squamate reptiles (lizards and snakes) based on 52 genes and 4162 species". Molecular Phylogenetics and Evolution. 94 (Pt B): 537–547. doi:10.1016/j.ympev.2015.10.009. PMID 26475614.

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[1] Han, D., K. Zhou, & A. M. Bauer. 2004. Phylogenetic relationships among gekkotan lizards inferred from c-mos nuclear DNA sequences and a new classification of the Gekkota. Biological Journal of the Linnean Society 83: 353– 368.

[2] Gamble, T., A. M. Bauer, E. Greenbaum, & T.R. Jackman. 2008. Evidence for Gondwanan vicariance in an ancient clade of gecko lizards. Journal of Biogeography 35: 88-104

[3] Pyron, R; Burbrink, Frank T; Wiens, John J (2013). "A phylogeny and revised classification of Squamata, including 4161 species of lizards and snakes". BMC Evolutionary Biology. 13: 93. Bibcode:2013BMCEE..13...93P. doi:10.1186/1471-2148-13-93. PMC 3682911. PMID 23627680.

[zheng_&_wiens_2016-4] Zheng, Yuchi; Wiens, John J. (2016). "Combining phylogenomic and supermatrix approaches, and a time-calibrated phylogeny for squamate reptiles (lizards and snakes) based on 52 genes and 4162 species". Molecular Phylogenetics and Evolution. 94 (Pt B): 537–547. doi:10.1016/j.ympev.2015.10.009. PMID 26475614.

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