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Ctenochelys

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Ctenochelys Temporal range: Late Cretaceous, 89–70 Ma PreꞒ Ꞓ O S D C P T J K Pg N

Ctenochelys stenoporus
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Order:	Testudines
Suborder:	Cryptodira
Superfamily:	Chelonioidea
Family:	†Ctenochelyidae
Genus:	†Ctenochelys Zangerl, 1953
Type species
†C. stenoporus
Species
†C. stenoporus †C. acris

Ctenochelys (meaning "comb tortoise") is an extinct genus of marine turtle (Cryptodira, Pancheloniidae), which existed during the Cretaceous period, and lived in the shallow waters of the Western Interior Seaway.^[1] Its fossils have been found in the Ripley Formation and Mooreville Chalk of central Alabama, United States.^[2] It was first named by C. H. Sternberg in 1904,^[3] and contains two species, C. stenoporus^[4] and C. acris.^[5]

Species

Ctenochelys stenoporus skeletons

Ctenochelys stenoporus is the type species. It was originally thought to be a species of Toxochelys; T. bauri, until Sternberg declared it a separate genus.^[3] The two genera are similar in carapaces.^[1]

Ctenochelys acris was first named by Zangerl in 1953 and is now thought to be one of the earliest ancestors of modern cheloniids.^[6]

Phylogeny

Ctenochelys in a cladogram from the analysis of Gentry et al., 2019:^[7]

Pan-Chelonioidea

†Angolachelonia

†Protostegidae

†Toxochelys latiremis

†Ctenochelyidae

†Asmodochelys parhami

	†Prionochelys matutina

	†Ctenochelys acris

	†Ctenochelys stenoporus

	†Peritresius ornatus

	†Peritresius martini

References

^ ^a ^b Kear BP, Lee MS (March 2006). "A primitive protostegid from Australia and early sea turtle evolution". Biol. Lett. 2 (1): 116–9. doi:10.1098/rsbl.2005.0406. PMC 1617175. PMID 17148342.
^ "Gary Kobylski, New Chair of MO–15 Board and State Conservationist in Alabama - Section "Things other than Dirt!"" (PDF). Charles Love, The Coastal Plainer. Fall 2005. Archived from the original (PDF) on 2009-03-20. Retrieved 2008-07-01.
^ ^a ^b "Marine Turtles". Mike Everhart. 2004. Archived from the original on 2013-12-25. Retrieved 2008-07-01.
^ Matzke AT (2007). "An almost complete juvenile specimen of the Cheloniid turtle Ctenochelys Stenoporus (Hay, 1905) From the Upper Cretaceous Niobrara formation of Kansas, USA". Palaeontology. 50 (3): 669–91. doi:10.1111/j.1475-4983.2007.00650.x.
^ Zangerl R (1953). "The vertebrate fauna of the Selma Formation of Alabama. Part IV. The turtles of the family Toxochelyidae". Fieldiana Geology Memoirs: 137–277.
^ Gentry AD (2016). "New material of the Late Cretaceous marine turtle Ctenochelys acris Zangerl, 1953 and a phylogenetic reassessment of the 'toxochelyid'-grade taxa". Journal of Systematic Palaeontology. 15 (8): 675–696. doi:10.1080/14772019.2016.1217087. S2CID 88758027.
^ Andrew D. Gentry; Jun A. Ebersole; Caitlin R. Kiernan (2019). "Asmodochelys parhami, a new fossil marine turtle from the Campanian Demopolis Chalk and the stratigraphic congruence of competing marine turtle phylogenies". Royal Society Open Science. 6 (12): Article ID 191950. Bibcode:2019RSOS....691950G. doi:10.1098/rsos.191950. PMC 6936288. PMID 31903219.

External links

Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Diapsida
Order: Testudines

Genus

Chelonioidea
(Sea turtles)

Cheloniidae	†Allopleuron Caretta †Carolinochelys Chelonia †Eochelone Eretmochelys †Gigantatypus †Glarichelys †Itilochelys Lepidochelys †Mexichelys †Miocaretta Natator †Pacifichelys †Syllomus †Tasbacka
Dermochelyidae	†Arabemys †Corsochelys †Cosmochelys Dermochelys †Eosphargis †Mesodermochelys †Psephophorus
	†Euclastes †Peritresius †Procolpochelys †Protosphargis †Puppigerus

Dermatemydidae	Dermatemys
Kinosternidae	Claudius †Hoplochelys Kinosternon Staurotypus Sternotherus

Emydidae	†Acherontemys Chrysemys Clemmys Deirochelys Emys Actinemys Emydoidea Glyptemys Graptemys Malaclemys Pseudemys Terrapene Trachemys †Wilburemys
Geoemydidae	Batagur †Banhxeochelys Cuora Cyclemys Geoclemys Geoemyda Hardella Heosemys Leucocephalon Malayemys Mauremys Melanochelys Morenia Notochelys Orlitia Pangshura Rhinoclemmys Sacalia Siebenrockiella Vijayachelys
Platysternidae	Platysternon
Testudinidae	Aldabrachelys Astrochelys Centrochelys Chelonoidis Chersina Cylindraspis †Cymatholcus †Floridemys Geochelone Gopherus †Hadrianus †Hesperotestudo Homopus Indotestudo Kinixys Malacochersus Manouria †Megalochelys †Oligopherus Psammobates Pyxis †Solitudo Stigmochelys †Stylemys Testudo

Carettochelyidae	†Allaeochelys †Anosteira Carettochelys
Trionychidae	Amyda Apalone †Axestemys Chitra Cyclanorbis Cycloderma †Drazinderetes Dogania †Gilmoremys †Hutchemys †Khunnuchelys Lissemys Nilssonia Palea †Palaeoamyda Pelochelys Pelodiscus Rafetus Trionyx
	†Basilochelys †Sinaspideretes

Chelydridae	Chelydra †Chelydrops †Chelydropsis †Emarginachelys †Macrocephalochelys Macrochelys †Planiplastron †Protochelydra
†Nanhsiungchelyidae	†Anomalochelys †Basilemys †Jiangxichelys
†Protostegidae	†Alienochelys †Archelon †Atlantochelys †Bouliachelys †Calcarichelys †Cratochelone †Desmatochelys †Iserosaurus †Notochelone †Ocepechelon †Pneumatoarthrus †Protostega †Rhinochelys †Santanachelys †Terlinguachelys
	†Adocus †Argillochelys †Bashuchelys †Ctenochelys †Prionochelys †Toxochelys

†Araripemydidae	†Araripemys
†Bothremydidae	†Araiochelys †Arenila †Azabbaremys †Bothremys †Cearachelys †Chedighaii †Chupacabrachelys †Eotaphrosphys †Foxemys †Galianemys †Ilatardia †Inaechelys †Itapecuruemys †Jainemys †Kinkonychelys †Kurmademys †Labrostochelys †Nigeremys †Phosphatochelys †Polysternon †Puentemys †Rosasia †Rhothonemys †Sankuchemys †Taphrosphys †Ummulisani †Zolhafah
Chelidae	Acanthochelys Chelodina Chelus Elseya Elusor Emydura Hydromedusa †Lomalatachelys Mesoclemmys Myuchelys Phrynops Platemys †Prochelidella Pseudemydura Rheodytes Rhinemys †Yaminuechelys
Pelomedusidae	Pelomedusa Pelusios
Podocnemididae	†Albertwoodemys †Bauruemys †Brontochelys †Caninemys †Carbonemys †Cerrejonemys †Cordichelys Erymnochelys †Lapparentemys †Latentemys Peltocephalus Podocnemis †Stupendemys
†Sahonachelyidae	†Sahonachelys †Sokatra

	†Caribemys †Caririemys †Tacuarembemys

Phylogenetic arrangement of turtles based on Turtles of the World 2017 Update: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status. † = extinct.

See also List of Testudines families

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